 He studied medicine at the Collège de France under Louis-Antoine Ranvier (1835-1922) and Louis-Charles Malassez (1842-1909) where he learned histological techniques and their correlation to other medical disciplines.He was "chef du laboratoire" at the medical clinic in the Hôtel-Dieu de Paris, and director of the histological laboratory at the École des Hautes Etudes.From 1925 to 1940 he was a professor at the Collège de France.