 He took a prominent role in the Estates General of 1560, the Colloquy of Poissy and the Colloquy of Saint-Germain in 1562.He participated in the papal conclaves of 1534, 1549, and 1559.He was the son of Jacques, seigneur de Tournon and Comte de Rousillon, and Jeanne de Polignac, daughter of Guillaume-Armand, Comte de Polignac.