 More to the south, it occurs in Spain and Portugal, although the species is restricted to karst springs in Central Portugal.In central Europe, this species has been recently introduced in the Austrian Danube, where it was first recorded in Tulln, Lower Austria in 2001.In the Czech Republic, it is now extinct in Bohemia; the only findings were in the Elbe river near Litoměřice in 1917, and the most recent findings of empty shells took place in 1943.