 They found two unrelated clades within Malagasy and Comoran samples of ""Miniopterus manavi"", neither of which was closely related to "M. newtoni" or to Tanzanian samples of "M. minor".During 2009, Steven Goodman and colleagues published two papers that found a total of five genetically and morphologically distinct species within "Miniopterus manavi" as defined by Peterson and colleagues (1995), up to four of which can be found in a single locality.In order to determine the true identity of "M. manavi", Goodman and Claude Maminirina obtained bats near the type locality of "M. manavi" (the site where the original material was collected, from which the species was described) for inclusion in the analysis; they also sequenced one of Thomas's original specimens.