 Among the five species they identified, "M. griveaudi" occurs on Grande Comore and Anjouan and in northern and western Madagascar; "M. aelleni" occurs on Anjouan and in northern and western Madagascar; "M. brachytragos" is found in northern Madagascar only; "M. mahafaliensis" is confined to the southwestern part of the island; and "M. manavi" itself is known only from the eastern edge of the Central Highlands.These five species are not each other's closest relatives according to analyses of cytochrome "b" sequences and their similarities reflect convergent evolution.Cytochrome "b" suggested that the closest relative of "M. manavi" is the slightly larger "M. petersoni" from southeastern Madagascar.