 It is, however, only convergent to the typical gudgeon subfamily (Gobioninae) and in fact among the closest relatives of the typical barbels ("Barbus" and the doubtfully distinct "Luciobarbus") – if it is not actually their closest relative.It is found in Bosnia and Herzegovina and Croatia, where it inhabits the karst.It occurs in some creeks in the Livanjsko field, in the Cetina, Krka and perhaps Zrmanja Rivers, and it was successfully introduced to Lake Blidin and Busko Blato.