 Each pinna is 0.5–1 cm (0.2–0.4 in) wide.The rachis of the leaf is often twisted 180 degrees, sometimes up to 360 degrees, though sometimes not at all.A male plant develops 1 to 4 male (or pollen-bearing) cones, which are fusiform (spindle-shaped), and measure 15–20 cm (6–8 in) high by 5–6 cm (2-2.4 in) wide.