 It was first discovered in forest litter of Greenbrier Cove, Great Smoky Mountains National Park, Tennessee.Another specimen was recorded near Robbinsville, Graham County, North Carolina, under a stone in a virgin cove hardwoods forest, southeast of the first find.About 30 years later, 22 new populations were found in the region, including habitats in southwestern Virginia, eastern Tennessee, western North Carolina, and northern Georgia.