 The morphological complexus of the various tissues he calls the "system of tissues"; and the physiological complexus of the various organs he calls the "system of organs".From his many accurate descriptions, it is evident that his researches extended to many animals, and particularly to insects.He innovated in comparative anatomy, with ideas not found in Malpighi, Morgagni, Leeuwenhoek, or Haller, the path-finders in microscopic anatomy.