 These other pathogens included "Colletotrichum acutatum" (ripe rot), "Greeneria uvicola" (bitter rot) and other fungi from the genera of "Aspergillus, Penicillium" and "Alternaria".Susceptibility of certain wine grape varieties were found to increase as the grapes reached higher Brix sugar levels.Further research in the 2000s, showed that some of the open canopy techniques used to prevent "Botrytis" grey rot may actually promote the development of some of these other types of rot.