 (2013), the members of a sample of Udege belong to haplogroup N9b (10/31 = 32.3%, probably descended from Jōmon people of northern Japan), haplogroup C4b1 (6/31 = 19.4%, also found among Evenks, Evens, Yakut, Dolgans, Buryat, Bargut, and Yukaghirs; three of the six Udege members of C4b1 belong to the C4b1f subclade marked by a T14153C mutation, which has only been observed in Udege to date), haplogroup M7a2a3 (5/31 = 16.1%, probably descended from Jōmon people, but also observed among Evenks in the basin of the Nyukzha River and among Buryat), haplogroup M8a1b (4/31 = 12.9%, related to Japanese haplogroup M8a1a), haplogroup M9a1a1a2 (3/31 = 9.7%, this subclade also has been found in a Nivkh individual and belongs to a mainly Japanese but also Korean, Khamnigan, Kalmyk, Chinese, and Tibetan branch of the M9a1a1 clade, which is widespread in East Asia and notably frequent among present-day Tibetans), haplogroup Y1a (2/31 = 6.5%, shared mainly with the Hezhen, other Amur Tungusic/Nani peoples, and Nivkhs), and C4a1a4a (1/31 = 3.2%, also observed among Tibetans, Bargut, Buryat, Kyrgyz, Altai Kizhi, Teleut, Shor, Yakut, Evenks, and Evens).The mitochondrial gene pool of the Ulchi, a Tungusic-speaking people who live in the lower basin of the Amur River, differs in detail both from the Ewenic peoples and from the Udege.According to Sukernik "et al."