 The female cones are open, with sporophylls 16–25 cm long, with two to four ovules per sporophyll.The lamina is lanceolate, with spined dentate margins and an apical spine.The male cones are solitary, ovoid, 15–22 cm long and 8–12 cm diameter, orange, the sporophylls 35–45 mm long with an upturned apical spine.