 He further divided the Gratioloideae into five tribes – Gratioleae, Angeloniaeae, Stemodieae, Limoselleae, and Lindernieae.He then divided the Gratioleae, with its 16 genera (and about 182 species) into three subtribes – Caprarinae, Dopatrinae, and Gratiolinae.The Gratiolinae had 10 genera (about 121 species) distributed through temperate and tropical America – "Bacopa" and "Mecardonia" (formerly "Herpestis"), "Amphianthus", "Gratiola", "Sophronanthe", "Benjaminia", "Scoparia", "Boelkea", "Maeviella", and "Braunblequetia".