 This has been observed in captivity, but it is unclear if wild beetles will engage in acts of hybridization.Certain species such as "D. grantii" and "D. hyllus" are believed to be sister species, while "D. tityus" is thought to be a sister taxon to the Central American "white Hercules" lineage.The intermediate species that bridges the "white Hercules" and the "giant Hercules" lineages is thought to be "D. maya".