 Since the great majority of caudate cells fire at very low rates, this activity almost always shows up as an increase in firing rate.Thus, eye movements begin with activation in the caudate nucleus, which inhibits the SNr via the direct GABAergic projections, which in turn disinhibits the SC.Extracellular dopamine in the basal ganglia has been linked to motivational states in rodents, with high levels being linked to satiated "euphoria", medium levels with seeking, and low with aversion.