 Several invertebrates have specialised behaviours or structures to feed on millipedes, including larval glowworm beetles, "Probolomyrmex" ants, chlamydephorid slugs, and predaceous dung beetles of the genera "Sceliages" and "Deltochilum".A large subfamily of assassin bugs, the Ectrichodiinae with over 600 species, has specialized in preying upon millipedes.Parasites of millipedes include nematodes, phaeomyiid flies, and acanthocephalans.