 The diplosegments of millipedes have evolved in conjunction with their burrowing habits, and nearly all millipedes adopt a mainly subterranean lifestyle.They use three main methods of burrowing; bulldozing, wedging and boring.Members of the orders Julida, Spirobolida and Spirostreptida, lower their heads and barge their way into the substrate, the collum being the portion of their exoskeleton that leads the way.