 Conversely, the later species, "T. carolomartini" and "T. yaucensis", lacking these marks, probably fed in deeper waters in a manner similar to manatees.The earlier species chewed with the jaws going up and down to mash food, whereas later species chewed with the jaws going front to back to grind it.The later species fed entirely on the seafloor, similar to sirenians (manatees and dugongs) and the extinct desmostylians, becoming specialist bottom feeders of seagrasses.