 The vast majority of species are found in the temperate regions, with about 257 species described from the Palaearctic region and 81 from the Nearctic.In the tropics, the genus is species-poor, with 36 species described from Indo-Australia, 13 from the Neotropics and 22 from the Afrotropical region.In 2012, a further 27 species were described from the Afrotropics.