 Similarly, the southern part of the inter-river system between the Bemarivo and Betsiboka Rivers, where little is known about the presence of the species, requires surveys.Groups of this species have a home range area amounting to 4-9 ha.A 2014 work in Ankarafantsika National Park suggests that population densities range from 5 to 100 ind/km²) and significant (negative) effects of road and forest edge and/or a (positive) effect of river proximity on densities.