 It occurs from the Betsiboka River, up to the Maevarano River, and are common in large area between these rivers.Extensive surveys of the species' distribution conducted in 2009, 2010, and 2011 led to the confirmation of its presence in most forest fragments between these rivers.Nevertheless, its eastern distribution limits are unclear, between the Sofia and Bemarivo Rivers, the species has twice been reported to be absent (Table 1).