 They were widespread in central, south-western and southern China, although their range is now highly fragmented.Their range spans the area from Qinghai east to Jiangsu and south to Sichuan, Guangxi and Guangdong; notably in the basins of the Yangtze, Yellow and Pearl Rivers.One is from the Pearl River basin (at least in Guangxi), two from the Yellow River basin, one from the Yangtze River basin (at least in Chongqing and Guizhou) and the final from the Qiantang River (at least in Anhui).