 But as is known today, pelicans are closer to storks (which have a hallux but no webbing) than to pseudotooth birds and evolved their fully webbed toes independently.With both a webbed and a hypotrophied hallux being apomorphic and paraphyletic, its absence in pseudotooth birds does not provide much information on their relationship.While giant Galloanserae were common and diverse in the Paleogene in particular, these (diatrymas and mihirungs) were flightless terrestrial birds; it is perhaps significant though that the only other "bone-toothed" birds known so far are the two species of the moa-nalo genus "Thambetochen", extinct giant flightless dabbling ducks from the Hawaiian Islands.