 The sternum had the deep and short shape typical of dynamic soarers, and bony outgrowths at the keel's forward margin securely anchored the furcula.The legs were proportionally short, the feet probably webbed and the hallux was vestigial or entirely absent; the tarsometatarsi (anklebones) resembled those of albatrosses while the arrangement of the front toes was more like in fulmars.Typical for pseudotooth birds was a second toe that attached a bit kneewards from the others and was noticeably angled outwards.