 He is also a Fellow of the Royal Society of Edinburgh, the Royal College of Pathologists, the Royal College of Surgeons of Edinburgh and a founder member of the Academy of Medical Sciences.He has won many international prizes for his work including the Joseph Steiner Prize, the Meyenburg Prize (1995), the Yvette Mayent Prize, the Paul Ehrlich Prize and the Cancer Research UK Lifetime Achievement Prize (2012).He has been awarded honorary degrees from the Universities of Toulouse, Birmingham, Aberdeen, Stirling, Abertay, Brno and Nottingham.