 After a big disturbance, the pattern of succession is from high light/low nitrogen towards high nitrogen/low light environment.Another article by Tilman that has received substantial citation is his 1994 "Ecology" article that encompasses the idea that large numbers of species can coexist in a small habitat even when they require the same limiting nutrient (such as nitrogen), as long as there is a tradeoff between the species.Basically it means that they can coexist because species that are good competitors are not as good at colonizing or reproducing.