 "Tapinella atrotomentosa" has a wound-activated defence mechanism whereby injured fruit bodies convert chemicals known as leucomentins into atromentin, butenolide, and the feeding deterrent osmundalactone.Atromentin had previously been identified as the pigment producing the brown colour of the cap, but was not characterized as a chemical defence compound until 1989.Other compounds produced by the fungus include the orange-yellow flavomentins and violet spiromentin pigments.