 Genetic analysis from a sample of tortoises from Volcan Wolf found 84 first-generation "C. nigra" hybrids, some less than 15 years old.The genetic diversity of these individuals is estimated to have required 38 "C. nigra" parents, many of which could still be alive on Isabela Island.The Pinta Island species ("C. abingdonii", now extinct) has been found to be most closely related to the species on the islands of San Cristóbal ("C. chathamensis") and Española ("C. hoodensis") which lie over 300 km (190 mi) away, rather than that on the neighbouring island of Isabela as previously assumed.