 He later refined this in 1838, placing those with universal veils into tribe "Volvaria", those with free gills and a discrete stipe into tribe "Pluteus", those with a "Tricholoma"-like shape into tribe "Entoloma", and those with a depressed cap and decurrent gills into tribe "Clitopilus".The small tribe "Leptonia" had convex fleshy membranaceous caps, the tribe "Nolanea", were slender fungi with bell-shaped caps and hollow stems, and lastly tribe "Eccilia" had umblicilate caps and adnate gills.Paul Kummer raised "Entoloma", "Nolanea", "Leptonia" and "Eccilia" to genus-level in 1871, though Lucien Quélet created a new genus "Rhodophyllus" uniting all those fungi with pinkish-red adnate or sinuate gills and angular spores, similar in scope to the original "Hyporhodius".