 In 1985, Roland McKay of the Queensland Museum published a comprehensive review of the family to resolve these relationships, although a number of species are still listed as doubtful, with McKay unable to locate the holotypes.Along with the review of previously described species, McKay described an additional seven species, a number of which he described as subspecies.After this 1985 paper, additional specimens came to light, proving that all the subspecies he had identified were individual species.