 A 2004 study determined that the amatoxin content of "G. marginata" varied from 78.17 to 243.61 µg/g of fresh weight.In this study, the amanitin amounts from certain "Galerina" specimens were higher than those from some "Amanita phalloides", a European fungus generally considered as the richest in amanitins.The authors suggest that "other parameters such as extrinsic factors (environmental conditions) and intrinsic factors (genetic properties) could contribute to the significant variance in amatoxin contents from different specimens."