 For example, in Holocene times, ancient pangolin ("Manis palaeojavanica"), panther ("Panthera" sp) and tapir ("Tapirus indicus") became locally extinct in Borneo.Of the 57 mammal species that were identified from archaeological remains in the Niah Caves, Sarawak, 13 were bats.Four of these were megachiropterans, "Pteropus vampyrus", "Rousettus amplexicaudatus", "Rousettus" sp and "Eonycteris spelaea", all of which remain extant species in Borneo and Peninsular Malaysia.