 Some important figures in the history of semiotics, are Charles Sanders Peirce, Roland Barthes, and Roman Jakobson.In modern times, its best-known figures include Umberto Eco, A. J. Greimas, Louis Hjelmslev, and Tullio De Mauro.Investigations on signs in non-human communications are subject to biosemiotics, a field founded in the late 20th century by Thomas Sebeok and Thure von Uexküll.