 Moreover, molecular analysis conducted separately by López-Fernández, "et al"., and De la Maza-Benignos, "et al"., did not support "H. tamasopoensis", "H. cyanoguttatus", "H. teporatus", and "H. carpintis" as distinct from each other.However, morphological and zoogeographical analysis by De la Maza-Benignos, "et al"., confirmed the validity of these four species, as well as differential evolutionary trends.Analysis also supported the separation of "Herichthys" into three distinct geomorphological groups + "H. minckleyi".