 The Gorteriinae contain two groups, one comprising "Berkheya", "Cullumia", "Cuspidia", "Didelta" and "Heterorhachis", the other one "Gorteria" and its close relatives of the genera "Gazania" and "Hirpicium".Recent comparison of homologous genes has seriously upset the delineation of the species within "Gorteria" and prompted the reassignment of "Hirpicium alienatum" and "H. integrifolium" to "Gorteria".According to this study, the species and their relationships are as expressed in the following tree.