 He specifically paired it with the genus "Histioneis" due to their morphological distinctiveness including what he described as a head-funnel and neck collar.Until DNA barcoding became accessible, specific demarcation in the genus was a significant challenge due to the variability in morphological traits (specifically the cingular lists).While beautiful figures were published in the early 20th century, often their scientific value was limited due to the incompleteness of morphological analyses.