 With his colleagues at the Jožef Stefan Institute, he was one of the first Slovene scientists to determine primary structure of those proteins, later focusing on their receptors and developing a hypothesis about the mechanism of their activity.In later years, he studied the molecular evolution of snake venom toxins.He co-discovered a novel mobile element of DNA and demonstrated horizontal gene transfer of this element between snakes and ruminants in evolutionary history which attracted significant attention of the scientific community.