 He performed his PhD studies at the Santa Fe Institute (SFI) in Santa Fe New Mexico, receiving his PhD from the Faculty of Biology at Utrecht University in 1999.This was followed by a year of post-doc studies at the SFI, and three years as a fellow at the Center of Studies in Physics and Biology at the Rockefeller University, New York.Since 2003 he is Professor of Computational Biology at the Biozentrum of the University of Basel, and group leader at the Swiss Institute of Bioinformatics since 2004.