 "Aplestosuchus" is distinguished form all other known Crocodyliformes by a unique combination of trait, including four autapomorphies.It possesses an autapomorphic nasal bone with dorsal midline crest, and an autapomorphic frontal longitudinal ridge that reaches the midline contact between the prefrontals.Its medial supratemporal rim is crest-shaped, and the ridge along the ectopterygoid-jugal suture is notched at its caudal portion.