 In 2004, Bowler and Marsh retained the species in "Niebla", while also indicating that secondary metabolites were usnic acid and unidentified triterpenes.The triterpene zeorin is frequently absent or in trace amounts in a number of species in the subgenus "Vermilacinia", whereas (-)-16 α-hydroxykaurane is found in all species taking into the correction.The absence of the diterpene has been reported in the subgenus "Cylindricaria" for two species in North America.