 The 2nd century AD geographer Pausanias also suggests that she is the daughter of Inachus and retells the story of Zeus falling in love with Io, the legendary wrath of Hera, and the metamorphosis by which Io becomes a beautiful white heifer.At another instant several generations later, Pausanias recounts another Io, descendant of Phoroneus, daughter of Iasus, who himself was the son of Argus and Ismene, the daughter of Asopus, or of Triopas and Sosis; Io's mother in the latter case was Leucane.Io's father was called Peiren in the "Catalogue of Women", and by Acusilaus, possibly a son of the elder Argus, also known as Peiras, Peiranthus or Peirasus.