 However, studies led by Oldrich Říčan in 2008 and 2016 suggested that several species within "Amphilophus" should be moved to the genus "Astatheros".Species proposed to be moved to "Astatheros" in 2008 were "A. alfari", "A. altifrons", "A. bussingi", "A. diquis", "A. longimanus", "A. macracanthus" (which would be the type species for "Astatheros"), "A. margaritifer", "A. rhytisma", "A. robertsoni" and "A. rostratus".Further genetic studies led Říčan to put "A. macracanthus" in "Astatheros", but to put "A. alfari", "A. altifrons", "A. bussingi", "A. diquis", "A. longimanus", "A. rhytisma", "A. robertsoni" and "A. rostratus" within the genus "Cribroheros".