 Unlike Carposinidae the mouthparts include "labial palps" with the second rather than third segment the longest.With other unusual structural characteristics of the caterpillar and adult, it could represent the sister lineage of all other extant members of this superfamily (Dugdale et al., 1999).The genus "Sisyroxena" from Madagascar is also notable for its unusual venation and wing scale sockets (Dugdale et al., 1999).