title: Dublin

text: There are many green-spaces around the city, and Dublin City Council manages over of parks. Public parks include the Phoenix Park, Herbert Park, St Stephen's Green, Saint Anne's Park and Bull Island. The Phoenix Park is about west of the city centre, north of the River Liffey. Its perimeter wall encloses , making it one of the largest walled city parks in Europe. It includes large areas of grassland and tree-lined avenues, and since the 17th century has been home to a herd of wild fallow deer. The residence of the President of Ireland (Áras an Uachtaráin), which was built in 1751, is located in the park. The park is also home to Dublin Zoo, Ashtown Castle, and the official residence of the United States Ambassador. Music concerts are also sometimes held in the park.


title: Dublin

text: St Stephen's Green is adjacent to one of Dublin's main shopping streets, Grafton Street, and to a shopping centre named after it, while on its surrounding streets are the offices of a number of public bodies.


title: Dublin

text: Saint Anne's Park is a public park and recreational facility, shared between Raheny and Clontarf, both suburbs on the Northside. The park, the second largest municipal park in Dublin, is part of a former estate assembled by members of the Guinness family, beginning with Benjamin Lee Guinness in 1835 (the largest municipal park is nearby (North) Bull Island, also shared between Clontarf and Raheny), featuring a 5 km beach.


title: Dublin

text: The Dublin region is the economic centre of Ireland, and was at the forefront of the country's economic expansion during the Celtic Tiger period. In 2009, Dublin was listed as the fourth richest city in the world by purchasing power and 10th richest by personal income. According to "Mercer's 2011 Worldwide Cost of Living Survey", Dublin is the 13th most expensive city in the European Union (down from 10th in 2010) and the 58th most expensive place to live in the world (down from 42nd in 2010). , approximately 874,400 people were employed in the Greater Dublin Area. Around 60% of people who are employed in Ireland's financial, ICT, and professional sectors are located in this area.


title: Dublin

text: A number of Dublin's traditional industries, such as food processing, textile manufacturing, brewing, and distilling have gradually declined, although Guinness has been brewed at the St. James's Gate Brewery since 1759. Economic improvements in the 1990s attracted a number of global pharmaceutical, information and communications technology companies to the city and Greater Dublin Area. Companies such as Microsoft, Google, Amazon, eBay, PayPal, Yahoo!, Facebook, Twitter, Accenture, TikTok and Pfizer now have European headquarters and/or operational bases in the city, with several located in enterprise clusters like the Digital Hub and Silicon Docks. The presence of these companies has driven economic expansion in the city and led to Dublin sometimes being referred to as the "Tech Capital of Europe".


title: Dublin

text: Financial services have also become important to the city since the establishment of Dublin's International Financial Services Centre in 1987. More than 500 operations are approved to trade under the IFSC programme. The centre is host to half of the world's top 50 banks and to half of the top 20 insurance companies. Many international firms have established major headquarters in the city, such as Citibank. The Irish Stock Exchange (ISEQ), Internet Neutral Exchange (INEX) and Irish Enterprise Exchange (IEX) are also located in Dublin. Dublin has been positioned as one of the main cities vying to host Financial Services companies hoping to retain access to the Eurozone after Brexit. The Celtic Tiger also led to a temporary boom in construction, with large redevelopment projects in the Dublin Docklands and Spencer Dock. Completed projects include the Convention Centre, the 3Arena, and the Bord Gáis Energy Theatre.


title: Dublin

text: In the second quarter of 2018, Dublin touched its lowest unemployment rate in a decade, when it fell down to 5.7% as reported by the Dublin Economic Monitor.


title: Dublin

text: The road network in Ireland is primarily focused on Dublin. The M50 motorway, a semi-ring road which runs around the south, west and north of the city, connects important national primary routes to the rest of the country. In 2008, the West-Link toll bridge was replaced by the eFlow barrier-free tolling system, with a three-tiered charge system based on electronic tags and car pre-registration.


title: Dublin

text: The first phase of a proposed eastern bypass for the city is the Dublin Port Tunnel, which officially opened in 2006 to mainly cater for heavy vehicles. The tunnel connects Dublin Port and the M1 motorway close to Dublin Airport. The city is also surrounded by an inner and outer orbital route. The inner orbital route runs approximately around the heart of the Georgian city and the outer orbital route runs primarily along the natural circle formed by Dublin's two canals, the Grand Canal and the Royal Canal, as well as the North and South Circular Roads.


title: Dublin

text: The 2016 TomTom Traffic Index ranked Dublin the 15th most congested city in the world and the 7th most congested in Europe.


title: Dublin

text: Dublin is served by a network of nearly 200 bus routes which cover the city and suburbs. The majority of these are provided by Dublin Bus, with a modest number having been transferred to Go Ahead Ireland in 2018. A number of smaller companies also operate. Fares are generally calculated on a stage system based on distance travelled. There are several different levels of fares, which apply on most services. A "Real Time Passenger Information" system was introduced at Dublin Bus bus stops in 2012 in which signs relay display the projected time of the next buses' arrival based on its GPS position. The National Transport Authority is responsible for integration of bus and rail services in Dublin and has been involved in introducing a pre-paid smart card, called a Leap card, which can be used on all of Dublin's public transport services.


title: Dublin

text: The 2011 Census showed that 5.9 percent of commuters in Dublin cycled. A 2013 report by Dublin City Council on traffic flows crossing the canals in and out of the city found that just under 10% of all traffic was made up of cyclists, representing an increase of 14.1% over 2012 and an 87.2% increase over 2006 levels and is attributed to measures, such as, the Dublinbikes bike rental scheme, the provision of cycle lanes, public awareness campaigns to promote cycling and the introduction of the 30 km/h city centre speed limit.


title: Dublin

text: Dublin City Council began installing cycle lanes and tracks throughout the city in the 1990s, and the city had over of specific on- and off-road tracks for cyclists. In 2011, the city was ranked 9th of major world cities on the "Copenhagenize Index of Bicycle-Friendly Cities". The same index showed a fall to 15th in 2015, and Dublin was outside the top 20 in 2017.


title: Dublin

text: Dublinbikes is a self-service bicycle rental scheme which has been in operation in Dublin since 2009. Sponsored by JCDecaux and Just Eat, the scheme consists of hundreds of unisex bicycles stationed at 44 terminals throughout the city centre. Users must make a subscription for either an annual Long Term Hire Card or purchase a three-day ticket. , Dublinbikes had over 66,000 long-term subscribers making over 2 million journeys per year.


title: Dublin

text: Heuston and Connolly stations are the two main railway termini in Dublin. Operated by Iarnród Éireann, the Dublin Suburban Rail network consists of five railway lines serving the Greater Dublin Area and commuter towns such as Drogheda and Dundalk in County Louth, Gorey in County Wexford, and extending as far as Portlaoise and once a day, Newry. One of the five lines is the electrified Dublin Area Rapid Transit (DART) line, which runs primarily along the coast of Dublin, comprising 31 stations, from Malahide and Howth southwards as far as Greystones in County Wicklow. Commuter rail operates on the other four lines using Irish Rail diesel multiple units. In 2013, passengers for DART and Dublin Suburban lines were 16 million and 11.7 million, respectively (around 75% of all Irish Rail passengers).


title: Dublin

text: Dublin once had an extensive system of trams but this was largely phased out by 1949. A new light rail system, often described as a tram system, the Luas, was launched in 2004, and is run by Transdev Ireland (under contract from Transport Infrastructure Ireland), carrying over 34 million passengers annually. The network consists of two interconnecting lines; the Red Line links the Docklands and city centre with the south-western suburbs of Tallaght and Saggart, while the Green Line connects northern inner city suburbs and the main city centre with suburbs to the south of the city including Sandyford and Brides Glen. Together these lines comprise a total 67 stations and of track. Construction of a 6 km extension to the Green Line, bringing it into the north of the city, commenced in June 2013 and was opened for passenger travel on 9 December 2017.


title: Dublin

text: A metro service is proposed under the name of Metrolink, and planned to run from Dublin's northside to Sandyford via Dublin Airport and St. Stephen's Green.


title: Dublin

text: Dublin Connolly is connected by bus to Dublin Port and ferries run by Irish Ferries and Stena Line to Holyhead for connecting trains on the North Wales Coast Line to Chester, Crewe and London Euston. Dublin Connolly to Dublin Port can be reached via Amiens Street, Dublin into Store Street or by Luas via Busáras where Dublin Bus operates services to the Ferry Terminal.


title: Dublin

text: Dublin Airport (owned and operated by DAA) is located north of Dublin city, near Swords in the administrative county of Fingal. The headquarters of Ireland's flag carrier Aer Lingus and regional airline CityJet are located there, and those of low-cost carrier Ryanair nearby. The airport offers a short and medium-haul network, domestic services to regional airports in Ireland, and long-haul services to the United States, Canada, the Middle East and Hong Kong. Dublin Airport is the 11th busiest in the European Union, and by far the busiest airport on the island of Ireland.


title: Dublin

text: In 2014, Dublin Airport was the 18th busiest airport in Europe, serving over 21 million passengers. By 2016 this increased to 27.9 million passengers passing through the airport, establishing an all-time record supported by growth in both short- and long-haul networks. In 2015 and 2016, transatlantic traffic grew, with 158 summer flights a week to North America, making it the sixth largest European hub for that route over the year. Transatlantic traffic was also the fastest-growing segment of the market for the airport in 2016, in which a 16% increase from 2015 brought the yearly number of passengers travelling between Dublin and North America to 2.9 million.


title: Dublin

text: From 2010 to 2016, Dublin Airport saw an increase of nearly 9.5 million passengers in its annual traffic, as the number of commercial aircraft movements has similarly followed a growth trend from 163,703 in 2013 to 191,233 in 2015.


title: Dublin

text: Dublin is also served by Weston Airport and other small facilities, by a range of helicopter operators, and the military and some State services use Casement Aerodrome nearby.


title: Dublin

text: Dublin is the largest centre of education in Ireland, and is home to four universities and a number of other higher education institutions. It was the European Capital of Science in 2012.


title: Dublin

text: The University of Dublin is the oldest university in Ireland, dating from the 16th century, and is located in the city centre. Its sole constituent college, Trinity College (TCD), was established by Royal Charter in 1592 under Elizabeth I. It was closed to Roman Catholics until 1793, and the Catholic hierarchy then banned Roman Catholics from attending until 1970. It is situated in the city centre, on College Green, and has over 18,000 students.


title: Dublin

text: The National University of Ireland (NUI) has its seat in Dublin, which is also the location of the associated "constituent university" of University College Dublin (UCD), which has over 30,000 students. Founded in 1854, it is now the largest university in Ireland. UCD's main campus is at Belfield, about from the city centre, in the southeastern suburbs.


title: Dublin

text: As of 2019, Dublin's principal, and Ireland's largest, institution for technological education and research, Dublin Institute of Technology (DIT), with origins in 1887, has merged with two major suburban third level institutions, Institute of Technology, Tallaght and Institute of Technology, Blanchardstown, to form Technological University Dublin, Ireland's second largest university by student population. The new university offers a wide range of courses in areas include engineering, architecture, the sciences, health, journalism, digital media, hospitality, business, art and design, music and the humanities programmes, and has three long-term campuses, at Grangegorman, Tallaght and Blanchardstown.


title: Dublin

text: Dublin City University (DCU), formerly the National Institute for Higher Education (NIHE) Dublin, offers courses in business, engineering, science, communication courses, languages and primary education. It has around 16,000 students, and its main campus is located about from the city centre, in the northern suburbs. Aside from the main Glasnevin Campus, the Drumcondra campuses includes the former St. Patrick's College of Education, Drumcondra now also hosting students from the nearby Mater Dei Institute of Education and students from the Church of Ireland College of Education at the DCU Campus at All Hallows College.


title: Dublin

text: The Royal College of Surgeons in Ireland (RCSI) conducts a medical school which is both a university (since 2019) and a recognised college of the NUI, and is situated at St. Stephen's Green in the city centre; there are also large medical schools within UCD and Trinity College. The National College of Art and Design (NCAD) provides education and research in art, design and media. The National College of Ireland (NCI) is also based in Dublin, as well as the Economic and Social Research Institute, a social science research institute, on Sir John Rogerson's Quay, and the Dublin Institute for Advanced Studies.


title: Dublin

text: The Institute of International and European Affairs is also in Dublin. Dublin Business School (DBS) is Ireland's largest private third level institution with over 9,000 students located on Aungier Street, and Griffith College Dublin has its main facility in Portobello. There are also smaller specialised colleges, including The Gaiety School of Acting. The Irish public administration and management training centre has its base in Dublin, the Institute of Public Administration provides a range of undergraduate and post graduate awards via the National University of Ireland and in some instances, Queen's University Belfast.


title: Dublin

text: Dublin is also home to the Royal Irish Academy, membership of which is considered Ireland's highest academic honour.


title: Dublin

text: The suburban town of Dún Laoghaire is home to the Dún Laoghaire Institute of Art, Design and Technology (IADT), which supports training and research in art, design, business, psychology and media technology. Dublin joined the UNESCO Global Network of Learning Cities in 2019.


title: Dublin

text: The City of Dublin is the area administered by Dublin City Council. The term "Dublin" is also used to the traditional County Dublin (the Dublin Region), which includes the city and the counties of Dún Laoghaire–Rathdown, Fingal and South Dublin. The population of the city of Dublin was 554,554 in the 2016 census, while the population of the urban area was 1,173,179. The County Dublin population was 1,273,069 and that of the Greater Dublin Area 1,904,806. The area's population is expanding rapidly, and it was estimated by the Central Statistics Office that it would reach 2.1 million by 2020.


title: Dublin

text: After World War Two, Italians were by far the largest immigrant group in both Dublin and Ireland and became synonymous with the catering and restaurant landscape. Since the late 1990s, Dublin has experienced a significant level of net immigration, with the greatest numbers coming from the European Union, especially the United Kingdom, Poland and Lithuania. There is also immigration from outside Europe, including from Brazil, India, the Philippines, China and Nigeria. Dublin is home to a greater proportion of newer arrivals than any other part of the country. Sixty percent of Ireland's Asian population lives in Dublin. Over 15% of Dublin's population was foreign-born in 2006.


title: Dublin

text: The capital attracts the largest proportion of non-Catholic migrants from other countries. Increased secularisation in Ireland has prompted a drop in regular Catholic church attendance in Dublin from over 90 percent in the mid-1970s down to 14 percent according to a 2011 survey and less than 2% in some areas


title: Dublin

text: According to the 2016 census, the population of Dublin was 86.2% white (including 862,381 white Irish [72.5%], 132,846 other white [13.2%] and 5,092 [0.5%] white Irish traveller), 2% black (23,892), and 4.6% Asian (46,626). Additionally, 2.7% (27,412) are from other ethnic or cultural background, while 4.9% (49,092) did not state their ethnicity. In terms of religion, 68.2% identified as Catholic, 12.7% as other stated religions, with 19.1% having no religion or no religion stated.


title: Dublin

text: , there were 1,367 families within the Dublin region living in homeless accommodation or other emergency housing.


title: Dublin

text: Dublin has a significant literary history, and produced many literary figures, including Nobel laureates William Butler Yeats, George Bernard Shaw and Samuel Beckett. Other influential writers and playwrights include Oscar Wilde, Jonathan Swift and the creator of Dracula, Bram Stoker. It is also the location of key and notable works of James Joyce, including "Ulysses", which is set in Dublin and includes much topical detail. "Dubliners" is a collection of short stories by Joyce about incidents and typical characters of the city during the early 20th century. Other renowned writers include J. M. Synge, Seán O'Casey, Brendan Behan, Maeve Binchy, John Banville and Roddy Doyle. Ireland's biggest libraries and literary museums are found in Dublin, including the National Print Museum of Ireland and National Library of Ireland. In July 2010, Dublin was named as a UNESCO City of Literature, joining Edinburgh, Melbourne and Iowa City with the permanent title.


title: Dublin

text: Handel's oratorio Messiah was first performed at Neal's Music Hall, in Fishamble Street, on 13 April 1742.


title: Dublin

text: There are several theatres within the city centre, and various well-known actors have emerged from the Dublin theatrical scene, including Noel Purcell, Michael Gambon, Brendan Gleeson, Stephen Rea, Colin Farrell, Colm Meaney and Gabriel Byrne. The best known theatres include the Gaiety, Abbey, Olympia, Gate, and Grand Canal. The Gaiety specialises in musical and operatic productions, and also opens its doors after the evening theatre production to host a variety of live music, dancing, and films. The Abbey was founded in 1904 by a group that included Yeats with the aim of promoting indigenous literary talent. It went on to provide a breakthrough for some of the city's most famous writers, such as Synge, Yeats himself and George Bernard Shaw. The Gate was founded in 1928 to promote European and American Avant Garde works. The Grand Canal Theatre is a newer 2,111 capacity theatre which opened in 2010 in the Grand Canal Dock area.


title: Dublin

text: Apart from being the focus of the country's literature and theatre, Dublin is also the focal point for much of Irish art and the Irish artistic scene. The Book of Kells, a world-famous manuscript produced by Celtic monks in AD 800 and an example of Insular art, is on display in Trinity College. The Chester Beatty Library houses a collection of manuscripts, miniature paintings, prints, drawings, rare books and decorative arts assembled by American mining millionaire (and honorary Irish citizen) Sir Alfred Chester Beatty (1875–1968). The collections date from 2700 BCE onwards and are drawn from Asia, the Middle East, North Africa and Europe.


title: Dublin

text: In addition public art galleries are found across the city and are free to visit, including the Irish Museum of Modern Art, the National Gallery, the Hugh Lane Municipal Gallery, the Douglas Hyde Gallery, the Project Arts Centre and the exhibition space of the Royal Hibernian Academy. Private galleries in Dublin include Green on Red Gallery, Kerlin Gallery, Kevin Kavanagh Gallery and Mother's Tankstation.


title: Dublin

text: Three branches of the National Museum of Ireland are located in Dublin: Archaeology in Kildare Street, Decorative Arts and History in Collins Barracks and Natural History in Merrion Street. Dublin is home to the National College of Art and Design, which dates from 1746, and Dublin Institute of Design, founded in 1991. Dublinia is a living history attraction showcasing the Viking and Medieval history of the city.


title: Dublin

text: Dublin has long had an 'underground' arts scene, with Temple Bar hosting artists in the 1980s, and spaces such as the Project Arts Centre acting as a hub for collectives and new exhibitions. "The Guardian" noted that Dublin's independent and underground arts flourished during the economic recession of c.2010. Dublin also has many dramatic, musical and operatic companies, including Festival Productions, Lyric Opera Productions, the Pioneers' Musical & Dramatic Society, Rathmines and Rathgar Musical Society, the Glasnevin Musical Society, Third Day Chorale, Second Age Theatre Company, Irish National Opera.


title: Dublin

text: Dublin was shortlisted to be World Design Capital 2014. Taoiseach Enda Kenny was quoted to say that Dublin "would be an ideal candidate to host the World Design Capital in 2014".


title: Dublin

text: In October 2021, Dublin was shortlisted for the European Commission's 2022 European Capital of Smart Tourism award along with Bordeaux, Copenhagen, Florence, Ljubljana, Palma de Mallorca and Valencia.


title: Dublin

text: Dublin has a vibrant nightlife and is reputedly one of Europe's most youthful cities, with an estimate of 50% of citizens being younger than 25. There are many pubs across the city centre, with the area around St. Stephen's Green and Grafton Street, especially Harcourt Street, Camden Street, Wexford Street and Leeson Street, the location of many nightclubs and pubs.


title: Dublin

text: The best known area for nightlife is Temple Bar, south of the River Liffey. The area has become popular among tourists, including stag and hen parties from the UK. It was developed as Dublin's cultural quarter and does retain this spirit as a centre for small arts productions, photographic and artists' studios, and in the form of street performers and small music venues; however, it has been criticised as overpriced, false and dirty by Lonely Planet. The areas around Leeson Street, Harcourt Street, South William Street and Camden/George's Street are popular nightlife spots for locals.


title: Dublin

text: Live music is played on streets and at venues throughout Dublin, and the city has produced several musicians and groups of international success, including The Dubliners, Thin Lizzy, The Boomtown Rats, U2, The Script, Sinéad O'Connor, Boyzone, Kodaline, Fontaines D.C. and Westlife. Dublin has several mid-range venues that host live music throughout the week, including Whelans and Vicar Street. The 3Arena venue in the Dublin Docklands plays host to visiting global performers.


title: Dublin

text: Dublin city centre is a popular shopping destination for both locals and tourists. The city has numerous shopping districts, particularly around Grafton Street and Henry Street. The city centre is also the location of large department stores, including Arnotts, Brown Thomas and (prior to its 2015 closure) Clerys.


title: Dublin

text: While the city has seen the loss of some traditional market sites, Moore Street remains one of the city's oldest trading districts. There has also been some growth in local farmers' markets and other markets. In 2007, Dublin Food Co-op relocated to a warehouse in The Liberties area, where it is home to market and community events. Suburban Dublin has several modern retail centres, including Dundrum Town Centre, Blanchardstown Centre, the Square in Tallaght, Liffey Valley Shopping Centre in Clondalkin, Omni Shopping Centre in Santry, Nutgrove Shopping Centre in Rathfarnham, Northside Shopping Centre in Coolock and Swords Pavilions in Swords.


title: Dublin

text: Dublin is the centre of both media and communications in Ireland, with many newspapers, radio stations, television stations and telephone companies based there. RTÉ is Ireland's national state broadcaster, and is based in Donnybrook. Fair City is RTÉ's soap opera, located in the fictional Dublin suburb of "Carraigstown".


title: Dublin

text: Virgin Media Television, eir Sport, MTV Ireland and Sky News are also based in the city. The headquarters of An Post and telecommunications companies such as Eir, as well as mobile operators Vodafone and 3 are all located there. Dublin is also the headquarters of national newspapers such as "The Irish Times" and "Irish Independent", as well as local newspapers such as "The Evening Herald".


title: Dublin

text: As well as being home to RTÉ Radio, Dublin also hosts the national radio networks Today FM and Newstalk, and local stations. Commercial radio stations based in the city include 4fm (94.9 MHz), Dublin's 98FM (98.1 MHz), Radio Nova 100FM (100.3 MHz), Q102 (102.2 MHz), SPIN 1038 (103.8 MHz), FM104 (104.4 MHz), Sunshine 106.8 (106.8 MHz). There are also numerous community and special interest stations, including Dublin City FM (103.2 MHz), Dublin South FM (93.9 MHz), Liffey Sound FM (96.4 MHz), Near FM (90.3 MHz), and Raidió Na Life (106.4 MHz).


title: Dublin

text: Croke Park is the largest sport stadium in Ireland. The headquarters of the Gaelic Athletic Association, it has a capacity of 82,300. It is the third-largest stadium in Europe after Nou Camp in Barcelona and Wembley Stadium in London. It hosts the premier Gaelic football and hurling games, international rules football and irregularly other sporting and non-sporting events including concerts. Muhammad Ali fought there in 1972 and it played host to the opening and closing ceremonies of the 2003 Special Olympics. It also has conference and banqueting facilities. There is a GAA Museum there and tours of the stadium are offered, including a rooftop walk of the stadium. During the redevelopment of Lansdowne Road, Croke Park played host to the Irish Rugby Union Team and Republic of Ireland national football team as well as hosting the Heineken Cup rugby 2008–09 semi-final between Munster and Leinster which set a world record attendance for a club rugby match. The Dublin GAA team plays most of their home league hurling games at Parnell Park.


title: Dublin

text: IRFU Stadium Lansdowne Road was laid out in 1874. This was the venue for home games of both the Irish Rugby Union Team and the Republic of Ireland national football team. A joint venture between the Irish Rugby Football Union, the FAI and the Government, saw it redeveloped into a new state-of-the-art 50,000 seat Aviva Stadium, which opened in May 2010. Aviva Stadium hosted the 2011 UEFA Europa League Final. Rugby union team Leinster Rugby play their competitive home games in the RDS Arena & the Aviva Stadium while Donnybrook Stadium hosts their friendlies and A games, Ireland A and Women, Leinster Schools and Youths and the home club games of All Ireland League clubs Old Wesley and Bective Rangers. County Dublin is home for 13 of the senior rugby union clubs in Ireland including 5 of the 10 sides in the top division 1A.


title: Dublin

text: County Dublin is home to six League of Ireland association football (soccer) clubs; Bohemians F.C., Shamrock Rovers, St Patrick's Athletic, University College Dublin, Shelbourne and Cabinteely. The first Irish side to reach the group stages of a European competition (2011–12 UEFA Europa League group stage) are Shamrock Rovers, who play at Tallaght Stadium in South Dublin. Bohemian F.C play at Dalymount Park, the oldest football stadium in the country, and home ground for the Ireland football team from 1904 to the 1970s. St Patrick's Athletic play at Richmond Park; University College Dublin at the UCD Bowl in Dún Laoghaire–Rathdown; and Shelbourne at Tolka Park. Tolka Park, Dalymount Park, UCD Bowl and Tallaght Stadium, along with the Carlisle Grounds in Bray, hosted all Group 3 games in the intermediary round of the 2011 UEFA Regions' Cup.


title: Dublin

text: Dublin has two ODI cricket grounds in Castle Avenue (Clontarf Cricket Club) and Malahide Cricket Club Ground. College Park has Test status and played host to Ireland's first Test cricket match, a women's match against Pakistan in 2000. The men's Irish cricket team also played their first Test match against Pakistan at Malahide Cricket Club Ground during 2018. Leinster Lightning play their home inter-provincial matches in Dublin at College Park.


title: Dublin

text: The Dublin Marathon has been run since 1980 at the end of October. The Women's Mini Marathon has been run since 1983 on the first Monday in June, which is also a bank holiday in Ireland. It is said to be the largest all female event of its kind in the world. The Great Ireland Run takes place in Dublin's Phoenix Park in mid-April.


title: Dublin

text: The Dublin area hosts greyhound racing at Shelbourne Park and horse racing at Leopardstown. The Dublin Horse Show takes place at the RDS, which hosted the Show Jumping World Championships in 1982. The national boxing arena is located in The National Stadium on the South Circular Road. The National Basketball Arena is located in Tallaght, is the home of the Irish basketball team, the venue for the basketball league finals, and has also hosted boxing and wrestling events. The National Aquatic Centre in Blanchardstown is Ireland's largest indoor water leisure facility. There are also Gaelic Handball, hockey and athletics stadia, most notably Morton Stadium in Santry, which held the athletics events of the 2003 Special Olympics.


title: Dublin

text: As of the 2018 Michelin Guide, five Dublin restaurants shared six Michelin stars – including Restaurant Patrick Guilbaud with two. Irish-born Kevin Thornton was awarded two Michelin stars in 2001 – though his restaurant, Thornton's, closed in 2016. The Dublin Institute of Technology commenced a bachelor's degree in culinary skills in 1999.


title: Dublin

text: Historically, Irish coffee houses and cafes were associated with those working in media. Since the beginning of the 21st century, with the growth of apartment living in the city, Dublin's cafés attracted younger patrons looking for an informal gathering place and an ad hoc office. Cafés became more popular in the city, and Irish-owned coffee chains like Java Republic, Insomnia, and O'Brien's Sandwich Bars now compete internationally. In 2008, Irish barista Stephen Morrissey won the title of World Barista Champion.


title: Dublin

text: Dublin was traditionally a city of two languages, English and Irish, a situation found also in the area around it, The Pale. The Irish of County Dublin represented the easternmost extension of a broad central dialect area which stretched between Leinster and Connacht, but had its own local characteristics. It may also have been influenced by the east Ulster dialect of County Meath and County Louth to the north.


title: Dublin

text: In the words of a 16th-century English administrator, William Gerard (1518–1581): "All Englishe, and the most part with delight, even in Dublin, speak Irishe". The Old English historian Richard Stanihurst (1547–1618) wrote as follows: "When their posteritie became not altogither so warie in keeping, as their ancestors were valiant in conquering, the Irish language was free dennized in the English Pale: this canker tooke such deep root, as the bodie that before was whole and sound, was by little and little festered, and in manner wholly putrified".


title: Dublin

text: English authorities of the Cromwellian period accepted the fact that Irish was widely spoken in the city and its surrounds. In 1655 several local dignitaries were ordered to oversee a lecture in Irish to be given in Dublin. In March 1656 a converted Catholic priest, Séamas Corcy, was appointed to preach in Irish at Bride's parish every Sunday, and was also ordered to preach at Drogheda and Athy. In 1657 the English colonists in Dublin presented a petition to the Municipal Council complaining that in Dublin itself "there is Irish commonly and usually spoken".


title: Dublin

text: In early 18th century Dublin, Irish was the language of a group of poets and scribes led by Seán and Tadhg Ó Neachtain. Scribal activity in Irish persisted in Dublin right through the 18th century. There were still native Irish speakers in County Dublin at the time of the 1851 census.


title: Dublin

text: Though the number of Irish speakers declined throughout Ireland in the 19th century, the end of the century saw a Gaelic revival, centred in Dublin and accompanied by renewed literary activity. This was the harbinger of a steady renewal of urban Irish, though with new characteristics of its own.


title: Dublin

text: Dublin now has many thousands of habitual Irish speakers, with the 2016 census showing that daily speakers (outside the education system) numbered 14,903. They form part of an urban Irish-speaking cohort which is generally better-educated than monoglot English speakers.


title: Dublin

text: The Dublin Irish-speaking cohort is supported by a number of Irish-medium schools. There are 12,950 students in the Dublin region attending 34 gaelscoileanna (Irish-language primary schools) and 10 gaelcholáistí (Irish-language secondary schools).


title: Dublin

text: Two Irish language radio stations, Raidió Na Life and RTÉ Raidió na Gaeltachta, have studios in the city, and the online station Raidió Rí-Rá broadcasts from studios in the city. A number of Irish language agencies are also located in the capital. Conradh na Gaeilge offers language classes, has a book shop and is a meeting place for different groups. The closest Gaeltacht to Dublin is the County Meath Gaeltacht of Ráth Cairn and Baile Ghib which is away.


title: Dublin

text: Dublin city council has an International Relations Unit, established in 2007. It works on hosting of international delegations, staff exchanges, international promotion of the city, twinning and partnerships, work with multi-city organisations such as Eurocities, economic partnerships and advice to other Council units.


title: Dublin

text: The city also has "friendship" or "co-operation agreements" with a number of other cities: Moscow (2009−) and St Petersburg (2010−) in Russia and Guadalajara in Mexico (2013−), and has previously proposed an agreement with Rio de Janeiro also. Previous agreements have included those with Mexico City (2014−2018), Tbilisi in Georgia (2014−2017) and Wuhan in China (2016−2019).


title: Neanderthal

text: Neanderthals (, also Homo neanderthalensis and erroneously Homo sapiens neanderthalensis), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While the "causes of Neanderthal disappearance about 40,000 years ago remain highly contested," demographic factors such as small population size, inbreeding and genetic drift, are considered probable factors. Other scholars have proposed competitive replacement, assimilation into the modern human genome (bred into extinction), great climatic change, disease, or a combination of these factors.


title: Neanderthal

text: It is unclear when the line of Neanderthals split from that of modern humans; studies have produced various intervals ranging from 315,000 to more than 800,000 years ago. The date of divergence of Neanderthals from their ancestor "H. heidelbergensis" is also unclear. The oldest potential Neanderthal bones date to 430,000 years ago, but the classification remains uncertain. Neanderthals are known from numerous fossils, especially from after 130,000 years ago. The type specimen, Neanderthal 1, was found in 1856 in the Neander Valley in present-day Germany. For much of the early 20th century, European researchers depicted Neanderthals as primitive, unintelligent, and brutish. Although knowledge and perception of them has markedly changed since then in the scientific community, the image of the unevolved caveman archetype remains prevalent in popular culture.


title: Neanderthal

text: Neanderthal technology was quite sophisticated. It includes the Mousterian stone-tool industry and ability to create fire and build cave hearths, make the adhesive birch bark tar, craft at least simple clothes similar to blankets and ponchos, weave, go seafaring through the Mediterranean, and make use of medicinal plants, as well as treat severe injuries, store food, and use various cooking techniques such as roasting, boiling, and smoking. Neanderthals made use of a wide array of food, mainly hoofed mammals, but also other megafauna, plants, small mammals, birds, and aquatic and marine resources. Although they were probably apex predators, they still competed with cave bears, cave lions, cave hyaenas, and other large predators. A number of examples of symbolic thought and Palaeolithic art have been inconclusively attributed to Neanderthals, namely possible ornaments made from bird claws and feathers or shells, collections of unusual objects including crystals and fossils, engravings, music production (possibly indicated by the Divje Babe flute), and Spanish cave paintings contentiously dated to before 65,000 years ago. Some claims of religious beliefs have been made. Neanderthals were likely capable of speech, possibly articulate, although the complexity of their language is not known.


title: Neanderthal

text: Compared with modern humans, Neanderthals had a more robust build and proportionally shorter limbs. Researchers often explain these features as adaptations to conserve heat in a cold climate, but they may also have been adaptations for sprinting in the warmer, forested landscape that Neanderthals often inhabited. Nonetheless, they had cold-specific adaptations, such as specialised body-fat storage and an enlarged nose to warm air (although the nose could have been caused by genetic drift). Average Neanderthal men stood around and women tall, similar to pre-industrial modern humans. The braincases of Neanderthal men and women averaged about and respectively, which is within the range of the values for modern humans. The Neanderthal skull was more elongated and had smaller parietal lobes and cerebellum, morphological traits used to assign specimens to species.


title: Neanderthal

text: The total population of Neanderthals remained low, proliferating weakly harmful gene variants, and precluding effective long-distance networks. Despite this, there is evidence of regional cultures and thus of regular communication between communities. They may have frequented caves, and moved between them seasonally. Neanderthals lived in a high-stress environment with high trauma rates, and about 80% died before the age of 40. The 2010 Neanderthal genome project's draft report presented evidence for interbreeding between Neanderthals and modern humans. It possibly occurred 316–219 thousand years ago, but more likely 100,000 years ago and again 65,000 years ago. Neanderthals also appear to have interbred with Denisovans, a different group of archaic humans, in Siberia. Around 1–4% of genomes of Eurasians, Australo-Melanesians, Native Americans, and North Africans is of Neanderthal ancestry, while the inhabitants of sub-Saharan Africa have only 0.3% of Neanderthal genes, save possible traces from early Sapiens-to-Neanderthal gene flow and/or more recent back-migration of Eurasians to Africa. In all, about 20% of distinctly Neanderthal gene variants survive today. Although many of the gene variants inherited from Neanderthals may have been detrimental and selected out, Neanderthal introgression appears to have affected the modern human immune system,


title: Neanderthal

text: and is also implicated in several other biological functions and structures, but a large portion appears to be non-coding DNA.


title: Neanderthal

text: Neanderthals are named after the Neandertal Valley in which the first identified specimen was found. The valley was spelled "Neanderthal" and the species was spelled "Neanderthaler" in German until the spelling reform of 1901. The spelling "Neandertal" for the species is occasionally seen in English, even in scientific publications, but the scientific name, "H. neanderthalensis", is always spelled with "th" according to the principle of priority. The vernacular name of the species in German is always "Neandertaler" ("inhabitant of the Neander Valley"), whereas "Neandertal" always refers to the valley. The valley itself was named after the late 17th century German theologian and hymn writer Joachim Neander, who often visited the area. His name in turn means ‘new man’, being a learned Graecization of the German surname "Neumann".


title: Neanderthal

text: "Neanderthal" can be pronounced using the (as in ) or the standard English pronunciation of "th" with the fricative /θ/ (as ).


title: Neanderthal

text: Neanderthal 1, the type specimen, was known as the "Neanderthal cranium" or "Neanderthal skull" in anthropological literature, and the individual reconstructed on the basis of the skull was occasionally called "the Neanderthal man". The binomial name "Homo neanderthalensis"—extending the name "Neanderthal man" from the individual specimen to the entire species, and formally recognising it as distinct from humans—was first proposed by Irish geologist William King in a paper read to the 33rd British Science Association in 1863. However, in 1864, he recommended that Neanderthals and modern humans be classified in different genera as he compared the Neanderthal braincase to that of a chimpanzee and argued that they were "incapable of moral and <nowiki>[</nowiki>theistic] conceptions".


title: Neanderthal

text: The first Neanderthal remains—Engis 2 (a skull)—were discovered in 1829 by Dutch naturalist Philippe-Charles Schmerling in the Grottes d'Engis, Belgium, but he thought it was a fossil modern human skull. In 1848, Gibraltar 1 from Forbes' Quarry was presented to the Gibraltar Scientific Society by their Secretary Lieutenant Edmund Henry Réné Flint, but was also thought to be a modern human skull. In 1856, local schoolteacher Johann Carl Fuhlrott recognised bones from Kleine Feldhofer Grotte in Neander Valley—Neanderthal 1 (the holotype specimen)—as distinct from modern humans, and gave them to German anthropologist Hermann Schaaffhausen to study in 1857. It comprised the cranium, thigh bones, right arm, left humerus and ulna, left ilium (hip bone), part of the right shoulder blade, and pieces of the ribs. Following Charles Darwin's "On the Origin of Species", Fuhlrott and Schaaffhausen argued the bones represented an ancient modern human form; Schaaffhausen, a social Darwinist, believed that humans linearly progressed from savage to civilised, and so concluded that Neanderthals were barbarous cave-dwellers. Fuhlrott and Schaaffhausen met opposition namely from the prolific pathologist Rudolf Virchow who argued against defining new species based on only a single find. In 1872, Virchow erroneously interpreted Neanderthal characteristics as evidence of senility, disease, and malformation instead of archaicness, which stalled Neanderthal research until the end of the century.


title: Neanderthal

text: By the early 20th century, numerous other Neanderthal discoveries were made, establishing "H. neanderthalensis" as a legitimate species. The most influential specimen was La Chapelle-aux-Saints 1 ("The Old Man") from La Chapelle-aux-Saints, France. French palaeontologist Marcellin Boule authored several publications, among the first to establish palaeontology as a science, detailing the specimen, but reconstructed him as slouching, ape-like, and only remotely related to modern humans. The 1912 'discovery' of Piltdown Man (a hoax), appearing much more similar to modern humans than Neanderthals, was used as evidence that multiple different and unrelated branches of primitive humans existed, and supported Boule's reconstruction of "H. neanderthalensis" as a far distant relative and an evolutionary dead-end. He fuelled the popular image of Neanderthals as barbarous, slouching, club-wielding primitives; this image was reproduced for several decades and popularised in science fiction works, such as the 1911 "The Quest for Fire" by J.-H. Rosny aîné and the 1927 "" by H. G. Wells where they are depicted as monsters. In 1911, Scottish anthropologist Arthur Keith reconstructed La Chapelle-aux-Saints 1 as an immediate precursor to modern humans, sitting next to a fire, producing tools, wearing a necklace, and having a more humanlike posture, but this failed to garner much scientific rapport, and Keith later abandoned his thesis in 1915.


title: Neanderthal

text: By the middle of the century, based on the exposure of Piltdown Man as a hoax as well as a reexamination of La Chapelle-aux-Saints 1 (who had osteoarthritis which caused slouching in life) and new discoveries, the scientific community began to rework its understanding of Neanderthals. Ideas such as Neanderthal behaviour, intelligence, and culture were being discussed, and a more humanlike image of them emerged. In 1939, American anthropologist Carleton Coon reconstructed a Neanderthal in a modern business suit and hat to emphasise that they would be, more or less, indistinguishable from modern humans had they survived into the present. William Golding's 1955 novel "The Inheritors" depicts Neanderthals as much more emotional and civilised. However, Boule's image continued to influence works until the 1960s. In modern-day, Neanderthal reconstructions are often very humanlike.


title: Neanderthal

text: Hybridisation between Neanderthals and early modern humans had been suggested early on, such as by English anthropologist Thomas Huxley in 1890, Danish ethnographer Hans Peder Steensby in 1907, and Coon in 1962. In the early 2000s, supposed hybrid specimens were discovered: Lagar Velho 1 and Muierii 1. However, similar anatomy could also have been caused by adapting to a similar environment rather than interbreeding. Neanderthal admixture was found to be present in modern populations in 2010 with the mapping of the first Neanderthal genome sequence. This was based on 3 specimens in Vindija Cave, Croatia, which contained almost 4% archaic DNA (allowing for near complete sequencing of the genome). However, there was approximately 1 error for every 200 letters (base pairs) based on the implausibly high mutation rate, probably due to the preservation of the sample. In 2012, British-American geneticist Graham Coop hypothesised that they instead found evidence of a different archaic human species interbreeding with modern humans, which was disproven in 2013 by the sequencing of a high-quality Neanderthal genome preserved in a toe bone from Denisova Cave, Siberia.


title: Neanderthal

text: Neanderthals are hominids in the genus "Homo", humans, and generally classified as a distinct species, "H. neanderthalensis", although sometimes as a subspecies of modern human as "H. sapiens neanderthalensis". This would necessitate the classification of modern humans as "H. sapiens sapiens".


title: Neanderthal

text: A large part of the controversy stems from the vagueness of the term "species", as it is generally used to distinguish two genetically isolated populations, but admixture between modern humans and Neanderthals is known to have occurred. However, the absence of Neanderthal-derived patrilineal Y-chromosome and matrilineal mitochondrial DNA (mtDNA) in modern humans, along with the underrepresentation of Neanderthal X chromosome DNA, could imply reduced fertility or frequent sterility of some hybrid crosses, representing a partial biological reproductive barrier between the groups, and therefore species distinction. In 2014 geneticist Svante Pääbo summarised the controversy, describing such "taxonomic wars" as unresolvable, "since there is no definition of species perfectly describing the case".


title: Neanderthal

text: Neanderthals are thought to have been more closely related to Denisovans than to modern humans. Likewise, Neanderthals and Denisovans share a more recent last common ancestor (LCA) than to modern humans, based on nuclear DNA (nDNA). However, Neanderthals and modern humans share a more recent mitochondrial LCA (observable by studying mtDNA). This likely resulted from an interbreeding event subsequent to the Neanderthal/Denisovan split which introduced another mtDNA line. This involved either introgression coming from an unknown archaic human into Denisovans, or introgression from an earlier unidentified modern human wave from Africa into Neanderthals.


title: Neanderthal

text: It is largely thought that "H. heidelbergensis" was the last common ancestor of Neanderthals, Denisovans, and modern humans before populations became isolated in Europe, Asia, and Africa, respectively. The taxonomic distinction between "H. heidelbergensis" and Neanderthals is mostly based on a fossil gap in Europe between 300 and 243 thousand years ago during marine isotope stage 8. "Neanderthals", by convention, are fossils which date to after this gap. However, 430,000-year old bones at Sima de los Huesos could represent early Neanderthals or a closely related group, and the 400,000-year old Aroeira 3 could represent a transitional phase. Ancestral and derived morphs could have lived concurrently. It is also possible that there was gene flow between Western Europe and Africa during the Middle Pleistocene, obscuring Neanderthal characteristics in such specimens, namely from Ceprano, Italy, and Sićevo Gorge, Serbia. The fossil record is much more complete from 130,000 years ago onwards, and specimens from this period make up the bulk of known Neanderthal skeletons. Dental remains from the Italian Visogliano and Fontana Ranuccio sites indicate that Neanderthal dental features had evolved by around 450–430 thousand years ago during the Middle Pleistocene.


title: Neanderthal

text: There are two main hypotheses regarding the evolution of Neanderthals following the Neanderthal/human split: two-phase and accretion. Two-phase argues that a single major environmental event—such as the Saale glaciation—caused European "H. heidelbergensis" to increase rapidly in body size and robustness, as well as undergoing a lengthening of the head (phase 1), which then led to other changes in skull anatomy (phase 2). However, Neanderthal anatomy may not have been driven entirely by adapting to cold weather. Accretion holds that Neanderthals slowly evolved over time from the ancestral "H. heidelbergensis", divided into four stages: early-pre-Neanderthals (MIS 12, Elster glaciation), pre-Neanderthals "sensu lato" (MIS 11–9, Holstein interglacial), early Neanderthals (MIS 7–5, Saale glaciation–Eemian), and classic Neanderthals "sensu stricto" (MIS 4–3, Würm glaciation).


title: Neanderthal

text: Numerous dates for the Neanderthal/human split have been suggested. The date of around 250,000 years ago cites ""H. helmei"" as being the last common ancestor (LCA), and the split is associated with the Levallois technique of making stone tools. The date of about 400,000 years ago uses "H. heidelbergensis" as the LCA. Estimates of 600,000 years ago assume that ""H. rhodesiensis"" was the LCA, which split off into modern human lineage and a Neanderthal/"H. heidelbergensis" lineage. 800,000 years ago has "H. antecessor" as the LCA, but different variations of this model would push the date back to 1 million years ago. However, a 2020 analysis of "H. antecessor" enamel proteomes suggests that "H. antecessor" is related but not a direct ancestor. DNA studies have yielded various results for the Neanderthal/human divergence time, such as 538–315, 553–321, 565–503, 654–475, 690–550, 765–550, 741–317, and 800–520 thousand years ago; and a dental analysis concluded before 800,000 years ago.


title: Neanderthal

text: Neanderthals and Denisovans are more closely related to each other than they are to modern humans, meaning the Neanderthal/Denisovan split occurred after their split with modern humans. Assuming a mutation rate of 1 × 10 or 0.5 × 10 per base pair (bp) per year, the Neanderthal/Denisovan split occurred around either 236–190 or 473–381 thousand years ago respectively. Using 1.1 × 10 per generation with a new generation every 29 years, the time is 744,000 years ago. Using 5 × 10 nucleotide sites per year, it is 616,000 years ago. Using the latter dates, the split had likely already occurred by the time hominins spread out across Europe, and unique Neanderthal features had begun evolving by 600–500 thousand years ago. Before splitting, Neanderthal/Denisovans (or "Neandersovans") migrating out of Africa into Europe apparently interbred with an unidentified "superarchaic" human species who were already present there; these superarchaics were the descendants of a very early migration out of Africa around 1.9 mya.


title: Neanderthal

text: Pre- and early Neanderthals, living before the Eemian interglacial (130,000 years ago), are poorly known and come mostly from Western European sites. From 130,000 years ago onwards, the quality of the fossil record increases dramatically with classic Neanderthals, who are recorded from Western, Central, Eastern, and Mediterranean Europe, as well as Southwest, Central, and Northern Asia up to the Altai Mountains in southern Siberia. Pre- and early Neanderthals, on the other hand, seem to have continuously occupied only France, Spain, and Italy, although some appear to have moved out of this "core-area" to form temporary settlements eastward (although without leaving Europe). Nonetheless, southwestern France has the highest density of sites for pre-, early, and classic Neanderthals.


title: Neanderthal

text: The southernmost find was recorded at Shuqba Cave, Levant; reports of Neanderthals from the North African Jebel Irhoud and Haua Fteah have been reidentified as "H. sapiens". Their easternmost presence is recorded at Denisova Cave, Siberia 85°E; the southeast Chinese Maba Man, a skull, shares several physical attributes with Neanderthals, although these may be the result of convergent evolution rather than Neanderthals extending their range to the Pacific Ocean. The northernmost bound is generally accepted to have been 55°N, with unambiguous sites known between 50–53°N, although this is difficult to assess because glacial advances destroy most human remains, and palaeoanthropologist Trine Kellberg Nielsen has argued that a lack of evidence of Southern Scandinavian occupation is (at least during the Eemian interglacial) due to the former explanation and a lack of research in the area. Middle Palaeolithic artefacts have been found up to 60°N on the Russian plains, but these are more likely attributed to modern humans. A 2017 study claimed the presence of "Homo" at the 130,000 year old Californian Cerutti Mastodon site in North America, but this is largely considered implausible.


title: Neanderthal

text: It is unknown how the rapidly fluctuating climate of the last glacial period (Dansgaard–Oeschger events) impacted Neanderthals, as warming periods would produce more favourable temperatures but encourage forest growth and deter megafauna, whereas frigid periods would produce the opposite. However, Neanderthals may have preferred a forested landscape. Populations may have peaked in cold but not extreme intervals, such as marine isotope stages 8 and 6 (respectively 300 and 191 thousand years ago during the Saale glaciation). It is possible their range expanded and contracted as the ice retreated and grew respectively to avoid permafrost areas, residing in certain refuge zones during glacial maxima. In 2021, Israeli anthropologist Israel Hershkovitz and colleagues suggested the 140 to 120 thousand years old Israeli Nesher Ramla remains, which feature a mix of Neanderthal and more ancient "H. erectus" traits, represent one such source population which recolonised Europe following a glacial period.


title: Neanderthal

text: Like modern humans, Neanderthals probably descended from a very small population with an effective population—the number of individuals who can bear or father children—of 3,000 to 12,000 approximately. However, Neanderthals maintained this very low population, proliferating weakly harmful genes due to the reduced effectivity of natural selection.


title: Neanderthal

text: Various studies, using mtDNA analysis, yield varying effective populations, such as about 1,000 to 5,000; 5,000 to 9,000 remaining constant; or 3,000 to 25,000 steadily increasing until 52,000 years ago before declining until extinction. Archaeological evidence suggests that there was a tenfold increase in the modern human population in Western Europe during the period of the Neanderthal/modern human transition, and Neanderthals may have been at a demographic disadvantage due to a lower fertility rate, a higher infant mortality rate, or a combination of the two. Estimates giving a total population in the higher tens of thousands are contested. A consistently low population may be explained in the context of the "Boserupian Trap": a population's carrying capacity is limited by the amount of food it can obtain, which in turn is limited by its technology. Innovation increases with population, but if the population is too low, innovation will not occur very rapidly and the population will remain low. This is consistent with the apparent 150,000 year stagnation in Neanderthal lithic technology.


title: Neanderthal

text: In a sample of 206 Neanderthals, based on the abundance of young and mature adults in comparison to other age demographics, about 80% of them above the age of 20 died before reaching 40. This high mortality rate was probably due to their high-stress environment. However, it has also been estimated that the age pyramids for Neanderthals and contemporary modern humans were the same. Infant mortality was estimated to have been very high for Neanderthals, about 43% in northern Eurasia.


title: Neanderthal

text: Neanderthals had more robust and stockier builds than typical modern humans, wider and barrel-shaped rib cages; wider pelvises; and proportionally shorter forearms and forelegs.


title: Neanderthal

text: Based on 45 Neanderthal long bones from 14 men and 7 women, the average height was for males and for females. For comparison, the average height of 28 males and 10 females Upper Palaeolithic humans is respectively and , although this decreases by nearer the end of the period based on 21 males and 15 females; and the average in the year 1900 was and , respectively. The fossil record shows that adult Neanderthals varied from about in height, although some may have grown much taller (73.8 to 184.8 cm based on footprint length and from 65.8 to 189.3 cm based on footprint width). For Neanderthal weight, samples of 26 specimens found an average of for males and for females. Using , the body mass index for Neanderthal males was calculated to be 26.9–28.2, which in modern humans correlates to being overweight. This indicates a very robust build. The Neanderthal LEPR gene concerned with storing fat and body heat production is similar to that of the woolly mammoth, and so was likely an adaptation for cold climate.


title: Neanderthal

text: The neck vertebrae of Neanderthals are longer and thicker than those of (most) modern humans, leading to stability, possibly due to different head shape and size. Although the Neanderthal thorax (where the ribcage is) was similar in size to modern humans, the longer and straighter ribs would have equated to a widened mid-lower thorax and stronger breathing in the lower thorax, which are indicative of a larger diaphragm and possibly greater lung capacity. The lung capacity of Kebara 2 was estimated to have been , compared to the average human capacity of for males and for females. The Neanderthal chest was also more pronounced (expanded front-to-back, or antero-posteriorly). The sacrum (where the pelvis connects to the spine) was more vertically inclined, and was placed lower in relation to the pelvis, causing the spine to be less curved (exhibit less lordosis) and to fold in on itself somewhat (to be invaginated). In modern populations, this condition affects just a proportion of the population, and is known as a lumbarized sacrum. Such modifications to the spine would have enhanced side-to-side (mediolateral) flexion, better supporting the wider lower thorax. It is claimed by some that this feature would be normal for all "Homo", even tropically-adapted "Homo ergaster" or "erectus", with the condition of a narrower thorax in most modern humans being a unique characteristic.


title: Neanderthal

text: Body proportions are usually cited as being "hyperarctic" as adaptations to the cold, because they are similar to those of human populations which developed in cold climates—the Neanderthal build is most similar to that of Inuit and Siberian Yupiks among modern humans—and shorter limbs result in higher retention of body heat. Nonetheless, Neanderthals from more temperate climates—such as Iberia—still retain the "hyperarctic" physique. In 2019, English anthropologist John Stewart and colleagues suggested Neanderthals instead were adapted for sprinting, because of evidence of Neanderthals preferring more warmer wooded areas over the colder mammoth steppe, and DNA analysis indicating a higher proportion of fast-twitch muscle fibres in Neanderthals than in modern humans. He explained their body proportions and greater muscle mass as adaptations to sprinting as opposed to the endurance-oriented modern human physique, as persistence hunting may only be effective in hot climates where the hunter can run prey to the point of heat exhaustion (hyperthermia). They had longer heel bones, reducing their ability for endurance running, and their shorter limbs would have reduced moment arm at the limbs, allowing for greater rotational force at the wrists and ankles without extra exertion of the rotating muscles at the elbows and knees by increasing the speed at which the muscles contracted, causing faster acceleration. In 1981, American palaeoanthropologist Erik Trinkaus made note of this alternate explanation, but considered it less likely.


title: Neanderthal

text: Neanderthals had a staightened chin, sloping forehead, and projecting nose, which also started somewhat higher on the face than in most modern humans. The Neanderthal skull is typically more elongated and less globular than that of most modern humans, and features much more of an occipital bun, or "chignon", a protrusion on the back of the skull, although it is within the range of variation for humans who have it. It is caused by the cranial base and temporal bones being placed higher and more towards the front of the skull, and a flatter skullcap. They also had larger eyes likely to adapt to the low-light environment.


title: Neanderthal

text: The projected Neanderthal nose and paranasal sinuses have generally been explained as having warmed air as it entered the lungs and retained moisture ("nasal radiator" hypothesis); If their noses were wider, it would differ to the generally narrowed shape in cold-adapted creatures, and that it would have been caused instead by genetic drift. Also, the sinuses reconstructed wide are not grossly large, being comparable in size to those of modern humans. However, if sinus size is not an important factor for breathing cold air, then the actual function would be unclear, so they may not be a good indicator of evolutionary pressures to evolve such a nose. Further, a computer reconstruction of the Neanderthal nose and predicted soft tissue patterns shows some similarities to those of modern Arctic peoples, potentially meaning the noses of both populations convergently evolved for breathing cold, dry air.


title: Neanderthal

text: Neanderthals featured a rather large jaw which was once cited as a response to a large bite force evidenced by heavy wearing of Neanderthal front teeth (the "anterior dental loading" hypothesis), but similar wearing trends are seen in contemporary humans. It could also have evolved to fit larger teeth in the jaw, which would better resist wear and abrasion, and the increased wear on the front teeth compared to the back teeth probably stems from repetitive use. Neanderthal dental wear patterns are most similar to those of modern Inuit. The incisors are large and shovel-shaped, and, compared to modern humans, there was an unusually high frequency of taurodontism, a condition where the molars are bulkier due to an enlarged pulp (tooth core). Taurodontism was once thought to have been a distinguishing characteristic of Neanderthals which lent some mechanical advantage or stemmed from repetitive use, but was more likely simply a product of genetic drift. The bite force of Neanderthals and modern humans is now thought to be about the same, about and in modern human males and females, respectively.


title: Neanderthal

text: The Neanderthal braincase averages 1,640 cc for males and 1,460 cc for females, which is significantly larger than the averages for all groups of extant humans; for example, modern European males average and females . For 28 modern human specimens from 190 to 25 thousand years ago, the average was about disregarding sex, and modern human brain size is suggested to have decreased since the Upper Palaeolithic. The largest Neanderthal brain, Amud 1, was calculated to be , one of the largest ever recorded in hominids. Both Neanderthal and human infants measure about .


title: Neanderthal

text: The Neanderthal skull is distinctively elongated, a characteristic shape known as "en bombe," with smaller parietal lobes and a smaller cerebellum, areas implicated in tool use, visuospatial integration, numeracy, creativity, higher-order conceptualization, muscle memory, and possibly language, attention, working memory, social abilities, and episodic memory. Neanderthal brains also have larger occipital lobes (relating to the classic occurrence of an occipital bun in Neanderthal skull anatomy), likely an adaptation to lower light conditions in Europe that implies internal differences in the proportionality of brain-internal regions, relative to "Homo sapiens," consistent with external measurements obtained with fossil skulls. Their brains also have larger temporal lobe poles, wider orbitofrontal cortex, and larger olfactory bulbs, suggesting potential differences in language comprehension and associations with emotions (temporal functions), decision making (the orbitofrontal cortex), and sense of smell (olfactory bulbs). Their brains also show different rates of brain growth and development. Such differences, while slight, would have been visible to natural selection and may underlie and explain differences in the material record in things like social behaviors, technological innovation, and artistic output.


title: Neanderthal

text: The lack of sunlight most likely led to the proliferation of lighter skin in Neanderthals, although it has been recently claimed that light skin in modern Europeans was not particularly prolific until perhaps the Bronze Age. Genetically, BNC2 was present in Neanderthals, which is associated with light skin colour; however, a second variation of BNC2 was also present, which in modern populations is associated with darker skin colour in the UK Biobank. DNA analysis of three Neanderthal females from southeastern Europe indicates that they had brown eyes, dark skin color, and brown hair; with one having red hair.


title: Neanderthal

text: In modern humans, skin and hair colour is regulated by the melanocyte-stimulating hormone—which increases the proportion of eumelanin (black pigment) to phaeomelanin (red pigment)—which is encoded by the MC1R gene. There are five known variants in modern humans of the gene which cause loss-of-function and are associated with light skin and hair colour, and another unknown variant in Neanderthals (the R307G variant) which could be associated with pale skin and red hair. The R307G variant was identified in a Neanderthal from Monti Lessini, Italy, and possibly Cueva del Sidrón, Spain. However, as in modern humans, red was probably not a very common hair colour because the variant is not present in many other sequenced Neanderthals.


title: Neanderthal

text: Maximum natural lifespan and the timing of adulthood, menopause, and gestation were most likely very similar to modern humans. However, it has been hypothesised, based on the growth rates of teeth and tooth enamel, that Neanderthals matured faster than modern humans although this is not backed up by age biomarkers. The main differences in maturation are the atlas bone in the neck as well as the middle thoracic vertebrae fused about 2 years later in Neanderthals than in modern humans, but this was more likely caused by a difference in anatomy rather than growth rate.


title: Neanderthal

text: Generally, models on Neanderthal caloric requirements report significantly higher intakes than those of modern humans because they typically assume Neanderthals had higher basal metabolic rates (BMRs) due to higher muscle mass, faster growth rate, and greater body heat production against the cold; and higher daily physical activity levels (PALs) due to greater daily travelling distances while foraging. However, using a high BMR and PAL, American archaeologist Bryan Hockett estimated that a pregnant Neanderthal would have consumed 5,500 calories per day, which would have necessitated a heavy reliance on big game meat; such a diet would have caused numerous deficiencies or nutrient poisonings, so he concluded that these are poorly warranted assumptions to make.


title: Neanderthal

text: Neanderthals may have been more active during dimmer light conditions rather than broad daylight because they lived in regions with reduced daytime hours, hunted large game (such predators typically hunt at night to enhance ambush tactics), and had large eyes and visual processing neural centres. Genetically, colour blindness (which may enhance mesopic vision) is typically correlated with northern-latitude populations, and the Neanderthals from Vindija Cave, Croatia, had some substitutions in the Opsin genes which could have influenced colour vision. However, the functional implications of these substitutions are inconclusive. Neanderthal-derived alleles near ASB1 and EXOC6 are associated with being an evening person, narcolepsy, and day-time napping.


title: Neanderthal

text: Neanderthals suffered a high rate of traumatic injury, with an estimated 79–94% of specimens showing evidence of healed major trauma, of which 37–52% were severely injured, and 13–19% injured before reaching adulthood. One extreme example is Shanidar 1, who shows signs of an amputation of the right arm likely due to a nonunion after breaking a bone in adolescence, osteomyelitis (a bone infection) on the left clavicle, an abnormal gait, vision problems in the left eye, and possible hearing loss (perhaps swimmer's ear). In 1995, Trinkaus estimated that about 80% succumbed to their injuries and died before reaching 40, and thus theorised that Neanderthals employed a risky hunting strategy ("rodeo rider" hypothesis). However, rates of cranial trauma are not significantly different between Neanderthals and Middle Palaeolithic modern humans (although Neanderthals seem to have had a higher mortality risk), there are few specimens of both Upper Palaeolithic modern humans and Neanderthals who died after the age of 40, and there are overall similar injury patterns between them. In 2012, Trinkaus concluded that Neanderthals instead injured themselves in the same way as contemporary humans, such as by interpersonal violence. A 2016 study looking at 124 Neanderthal specimens argued that high trauma rates were instead caused by animal attacks, and found that about 36% of the sample were victims of bear attacks, 21% big cat attacks, and 17% wolf attacks (totalling 92 positive cases, 74%). There were no cases of hyaena attacks, although hyaenas still nonetheless probably attacked Neanderthals, at least opportunistically. Such intense predation probably stemmed from common confrontations due to competition over food and cave space, and from Neanderthals hunting these carnivores.


title: Neanderthal

text: Low population caused a low genetic diversity and probably inbreeding, which reduced the population's ability to filter out harmful mutations (inbreeding depression). However, it is unknown how this affected a single Neanderthal's genetic burden and, thus, if this caused a higher rate of birth defects than in modern humans. It is known, however, that the 13 inhabitants of Sidrón Cave collectively exhibited 17 different birth defects likely due to inbreeding or recessive disorders. Likely due to advanced age (60s or 70s), La Chapelle-aux-Saints 1 had signs of Baastrup's disease, affecting the spine, and osteoarthritis. Shanidar 1, who likely died at about 30 or 40, was diagnosed with the most ancient case of diffuse idiopathic skeletal hyperostosis (DISH), a degenerative disease which can restrict movement, which, if correct, would indicate a moderately high incident rate for older Neanderthals.


title: Neanderthal

text: Neanderthals were subject to several infectious diseases and parasites. Modern humans likely transmitted diseases to them; one possible candidate is the stomach bacteria "Helicobacter pylori". The modern human papillomavirus variant 16A may descend from Neanderthal introgression. A Neanderthal at Cueva del Sidrón, Spain, shows evidence of a gastrointestinal "Enterocytozoon bieneusi" infection. The leg bones of the French La Ferrassie 1 feature lesions that are consistent with periostitis—inflammation of the tissue enveloping the bone—likely a result of hypertrophic osteoarthropathy, which is primarily caused by a chest infection or lung cancer. Neanderthals had a lower cavity rate than modern humans, despite some populations consuming typically cavity-causing foods in great quantity, which could indicate a lack of cavity-causing oral bacteria, namely "Streptococcus mutans".


title: Neanderthal

text: Two 250,000-year-old Neanderthaloid children from Payré, France, present the earliest known cases of lead exposure of any hominin. They were exposed on two distinct occasions either by eating or drinking contaminated food or water, or inhaling lead-laced smoke from a fire. There are two lead mines within of the site.


title: Neanderthal

text: Neanderthals likely lived in more sparsely distributed groups than contemporary modern humans, but group size is thought to have averaged 10 to 30 individuals, similar to modern hunter-gatherers. Reliable evidence of Neanderthal group composition comes from Cueva del Sidrón, Spain, and the footprints at Le Rozel, France: the former shows 7 adults, 3 adolescents, 2 juveniles, and an infant; whereas the latter, based on footprint size, shows a group of 10 to 13 members where juveniles and adolescents made up 90%.


title: Neanderthal

text: A Neanderthal child's teeth analysed in 2018 showed it was weaned after 2.5 years, similar to modern hunter gatherers, and was born in the spring, which is consistent with modern humans and other mammals whose birth cycles coincide with environmental cycles. Indicated from various ailments resulting from high stress at a low age, such as stunted growth, British archaeologist Paul Pettitt hypothesised that children of both sexes were put to work directly after weaning; and Trinkaus said that, upon reaching adolescence, an individual may have been expected to join in hunting large and dangerous game. However, the bone trauma is comparable to modern Inuit, which could suggest a similar childhood between Neanderthals and contemporary modern humans. Further, such stunting may have also resulted from harsh winters and bouts of low food resources.


title: Neanderthal

text: Sites showing evidence of no more than three individuals may have represented nuclear families or temporary camping sites for special task groups (such as a hunting party). Bands likely moved between certain caves depending on the season, indicated by remains of seasonal materials such as certain foods, and returned to the same locations generation after generation. Some sites may have been used for over 100 years. Cave bears may have greatly competed with Neanderthals for cave space, and there is a decline in cave bear populations starting 50,000 years ago onwards (although their extinction occurred well after Neanderthals had died out). Although Neanderthals are generally considered to have been cave dwellers, with 'home base' being a cave, open-air settlements near contemporaneously inhabited cave systems in the Levant could indicate mobility between cave and open-air bases in this area. Evidence for long-term open-air settlements is known from the 'Ein Qashish site in Israel, and Moldova I in Ukraine. Although Neanderthals appear to have had the ability to inhabit a range of environments—including plains and plateaux—open-air Neanderthals sites are generally interpreted as having been used as slaughtering and butchering grounds rather than living spaces.


title: Neanderthal

text: In 2022, remains of the first-known Neanderthal family (six adults and five children) were excavated from Chagyrskaya Cave in the Altai Mountains of southern Siberia in Russia. The family, which included a father, a daughter, and what appear to be cousins, most likely died together, presumably from starvation.


title: Neanderthal

text: Canadian ethnoarchaeologist Brian Hayden calculated a self-sustaining population that avoids inbreeding to consist of about 450–500 individuals, which would necessitate these bands to interact with 8–53 other bands, but more likely the larger estimate given low population density. Analysis of the mtDNA of the Neanderthals of Cueva del Sidrón, Spain, showed that the three adult men belonged to the same maternal lineage, while the three adult women belonged to different ones. This suggests a patrilocal residence (that a woman moved out of her group to live with her husband). However, the DNA of a Neanderthal from Denisova Cave, Russia, shows that she had an inbreeding coefficient of (her parents were either half-siblings with a common mother, double first cousins, an uncle and niece or aunt and nephew, or a grandfather and granddaughter or grandmother and grandson) and the inhabitants of Cueva del Sidrón show several defects, which may have been caused by inbreeding or recessive disorders.


title: Neanderthal

text: Considering most Neanderthal artifacts were sourced no more than from the main settlement, Hayden considered it unlikely these bands interacted very often, and mapping of the Neanderthal brain and their small group size and population density could indicate that they had a reduced ability for inter-group interaction and trade. However, a few Neanderthal artefacts in a settlement could have originated 20, 30, 100, and 300 km (12.5, 18.5, 60, and 185 mi) away. Based on this, Hayden also speculated that macro-bands formed which functioned much like those of the low-density hunter-gatherer societies of the Western Desert of Australia. Macro-bands collectively encompass , with each band claiming , maintaining strong alliances for mating networks or to cope with leaner times and enemies. Similarly, British anthropologist Eiluned Pearce and Cypriot archaeologist Theodora Moutsiou speculated that Neanderthals were possibly capable of forming geographically expansive ethnolinguistic tribes encompassing upwards of 800 people, based on the transport of obsidian up to from the source compared to trends seen in obsidian transfer distance and tribe size in modern hunter-gatherers. However, according to their model Neanderthals would not have been as efficient at maintaining long-distance networks as modern humans, probably due to a significantly lower population. Hayden noted an apparent cemetery of six or seven individuals at La Ferrassie, France, which, in modern humans, is typically used as evidence of a corporate group which maintained a distinct social identity and controlled some resource, trading, manufacturing, and so on. La Ferrassie is also located in one of the richest animal-migration routes of Pleistocene Europe.


title: Neanderthal

text: Genetic analysis indicates there were at least 3 distinct geographical groups—Western Europe, the Mediterranean coast, and east of the Caucasus—with some migration among these regions. Post-Eemian Western European Mousterian lithics can also be broadly grouped into 3 distinct macro-regions: Acheulean-tradition Mousterian in the southwest, Micoquien in the northeast, and Mousterian with bifacial tools (MBT) in between the former two. MBT may actually represent the interactions and fusion of the two different cultures. Southern Neanderthals exhibit regional anatomical differences from northern counterparts: a less protrusive jaw, a shorter gap behind the molars, and a vertically higher jawbone. These all instead suggest Neanderthal communities regularly interacted with neighbouring communities within a region, but not as often beyond.


title: Neanderthal

text: Nonetheless, over long periods of time, there is evidence of large-scale cross-continental migration. Early specimens from Mezmaiskaya Cave in the Caucasus and Denisova Cave in the Siberian Altai Mountains differ genetically from those found in Western Europe, whereas later specimens from these caves both have genetic profiles more similar to Western European Neanderthal specimens than to the earlier specimens from the same locations, suggesting long-range migration and population replacement over time. Similarly, artefacts and DNA from Chagyrskaya and Okladnikov Caves, also in the Altai Mountains, resemble those of eastern European Neanderthal sites about away more than they do artefacts and DNA of the older Neanderthals from Denisova Cave, suggesting two distinct migration events into Siberia. Neanderthals seem to have suffered a major population decline during MIS 4 (71–57 thousand years ago), and the distribution of the Micoquian tradition could indicate that Central Europe and the Caucasus were repopulated by communities from a refuge zone either in eastern France or Hungary (the fringes of the Micoquian tradition) who dispersed along the rivers Prut and Dniester.


title: Neanderthal

text: There is also evidence of inter-group conflict: a skeleton from La Roche à Pierrot, France, showing a healed fracture on top of the skull apparently caused by a deep blade wound, and another from Shanadir Cave, Iraq, found to have a rib lesion characteristic of projectile weapon injuries.


title: Neanderthal

text: It is sometimes suggested, since they were hunters of challenging big game and lived in small groups, there was no sexual division of labour as seen in modern hunter-gatherer societies. That is, men, women, and children all had to be involved in hunting, instead of men hunting with women and children foraging. However, with modern hunter-gatherers, the higher the meat dependency, the higher the division of labour. Further, tooth-wearing patterns in Neanderthal men and women suggest they commonly used their teeth for carrying items, but men exhibit more wearing on the upper teeth, and women the lower, suggesting some cultural differences in tasks.


title: Neanderthal

text: It is controversially proposed that some Neanderthals wore decorative clothing or jewellery—such as a leopard skin or raptor feathers—to display elevated status in the group. Hayden postulated that the small number of Neanderthal graves found was because only high-ranking members would receive an elaborate burial, as is the case for some modern hunter-gatherers. Trinkaus suggested that elderly Neanderthals were given special burial rites for lasting so long given the high mortality rates. Alternatively, many more Neanderthals may have received burials, but the graves were infiltrated and destroyed by bears. Given that 20 graves of Neanderthals aged under 4 have been found—over a third of all known graves—deceased children may have received greater care during burial than other age demographics.


title: Neanderthal

text: Looking at Neanderthal skeletons recovered from several natural rock shelters, Trinkaus said that, although Neanderthals were recorded as bearing several trauma-related injuries, none of them had significant trauma to the legs that would debilitate movement. He suggested that self worth in Neanderthal culture derived from contributing food to the group; a debilitating injury would remove this self-worth and result in near-immediate death, and individuals who could not keep up with the group while moving from cave to cave were left behind. However, there are examples of individuals with highly debilitating injuries being nursed for several years, and caring for the most vulnerable within the community dates even further back to "H. heidelbergensis". Especially given the high trauma rates, it is possible that such an altruistic strategy ensured their survival as a species for so long.


title: Neanderthal

text: Neanderthals were once thought of as scavengers, but are now considered to have been apex predators. In 1980, it was hypothesised that two piles of mammoth skulls at La Cotte de St Brelade, Jersey, at the base of a gulley were evidence of mammoth drive hunting (causing them to stampede off a ledge), but this is contested. Living in a forested environment, Neanderthals were likely ambush hunters, getting close to and attacking their target—a prime adult—in a short burst of speed, thrusting in a spear at close quarters. Younger or wounded animals may have been hunted using traps, projectiles, or pursuit. Nonetheless, they were able to adapt to a variety of habitats. They appear to have eaten predominantly what was abundant within their immediate surroundings, with steppe-dwelling communities (generally outside of the Mediterranean) subsisting almost entirely on meat from large game, forest-dwelling communities consuming a wide array of plants and smaller animals, and waterside communities gathering aquatic resources. Contemporary humans, in contrast, seem to have used more complex food extraction strategies and generally had a more diverse diet. Nonetheless, Neanderthals still would have had to have eaten a varied enough diet to prevent nutrient deficiencies and protein poisoning, especially in the winter when they presumably ate mostly lean meat. Any food with high contents of other essential nutrients not provided by lean meat would have been vital components of their diet, such as fat-rich brains, carbohydrate-rich and abundant underground storage organs (including roots and tubers), or, like modern Inuit, the stomach contents of herbivorous prey items.


title: Neanderthal

text: For meat, they appear to have fed predominantly on hoofed mammals, namely red deer and reindeer as these two were the most abundant game, but also on other Pleistocene megafauna such as ibex, wild boar, aurochs, mammoth, straight-tusked elephant, woolly rhinoceros, and so on. There is evidence of directed cave and brown bear hunting both in and out of hibernation, as well as butchering. Analysis of Neanderthal bone collagen from Vindija Cave, Croatia, shows nearly all of their protein needs derived from animal meat. Some caves show evidence of regular rabbit and tortoise consumption. At Gibraltar sites, there are remains of 143 different bird species, many ground-dwelling such as the common quail, corn crake, woodlark, and crested lark. Neanderthals also exploited marine resources on the Iberian, Italian, and Peloponnesian Peninsulas, where they waded or dived for shellfish, as early as 150,000 years ago at Cueva Bajondillo, Spain, similar to the fishing record of modern humans. At Vanguard Cave, Gibraltar, the inhabitants consumed Mediterranean monk seal, short-beaked common dolphin, common bottlenose dolphin, Atlantic bluefin tuna, sea bream, and purple sea urchin; and at Gruta da Figueira Brava, Portugal, there is evidence of large-scale harvest of shellfish, crabs, and fish. Evidence of freshwater fishing was found in Grotte di Castelcivita, Italy, for trout, chub, and eel; Abri du Maras, France, for chub and European perch; Payré, France; and Kudaro Cave, Russia, for Black Sea salmon.


title: Neanderthal

text: Edible plant and mushroom remains are recorded from several caves. Neanderthals from Cueva del Sidrón, Spain, based on dental tartar, likely had a meatless diet of mushrooms, pine nuts, and moss, indicating they were forest foragers. Remnants from Amud Cave, Israel, indicates a diet of figs, palm tree fruits, and various cereals and edible grasses. Several bone traumas in the leg joints could possibly suggest habitual squatting, which, if the case, was likely done while gathering food. Dental tartar from Grotte de Spy, Belgium, indicates the inhabitants had a meat-heavy diet including woolly rhinoceros and mouflon sheep, while also regularly consuming mushrooms. Neanderthal faecal matter from El Salt, Spain, dated to 50,000 years ago—the oldest human faecal matter remains recorded—show a diet mainly of meat but with a significant component of plants. Evidence of cooked plant foods—mainly legumes and, to a far lesser extent, acorns—was discovered in Kebara Cave, Israel, with its inhabitants possibly gathering plants in spring and fall and hunting in all seasons except fall, although the cave was probably abandoned in late summer to early fall. At Shanidar Cave, Iraq, Neanderthals collected plants with various harvest seasons, indicating they scheduled returns to the area to harvest certain plants, and that they had complex food-gathering behaviours for both meat and plants.


title: Neanderthal

text: Neanderthals probably could employ a wide range of cooking techniques, such as roasting, and they may have been able to heat up or boil soup, stew, or animal stock. The abundance of animal bone fragments at settlements may indicate the making of fat stocks from boiling bone marrow, possibly taken from animals that had already died of starvation. These methods would have substantially increased fat consumption, which was a major nutritional requirement of communities with low carbohydrate and high protein intake. Neanderthal tooth size had a decreasing trend after 100,000 years ago, which could indicate an increased dependence on cooking or the advent of boiling, a technique that would have softened food.


title: Neanderthal

text: At Cueva del Sidrón, Spain, Neanderthals likely cooked and possibly smoked food, as well as used certain plants—such as yarrow and camomile—as flavouring, although these plants may have instead been used for their medicinal properties. At Gorham's Cave, Gibraltar, Neanderthals may have been roasting pinecones to access pine nuts.


title: Neanderthal

text: At Grotte du Lazaret, France, a total of 23 red deer, 6 ibexes, 3 aurochs, and 1 roe deer appear to have been hunted in a single autumn hunting season, when strong male and female deer herds would group together for rut. The entire carcasses seem to have been transported to the cave and then butchered. Because this is such a large amount of food to consume before spoilage, it is possible these Neanderthals were curing and preserving it before winter set in. At 160,000 years old, it is the oldest potential evidence of food storage. The great quantities of meat and fat which could have been gathered in general from typical prey items (namely mammoths) could also indicate food storage capability. With shellfish, Neanderthals needed to eat, cook, or in some manner preserve them soon after collection, as shellfish spoils very quickly. At Cueva de los Aviones, Spain, the remains of edible, algae eating shellfish associated with the alga "Jania rubens" could indicate that, like some modern hunter gatherer societies, harvested shellfish were held in water-soaked algae to keep them alive and fresh until consumption.


title: Neanderthal

text: Competition from large Ice Age predators was rather high. Cave lions likely targeted horses, large deer and wild cattle; and leopards primarily reindeer and roe deer; which heavily overlapped with Neanderthal diet. To defend a kill against such ferocious predators, Neanderthals may have engaged in a group display of yelling, arm waving, or stone throwing; or quickly gathered meat and abandoned the kill. However, at Grotte de Spy, Belgium, the remains of wolves, cave lions, and cave bears—which were all major predators of the time—indicate Neanderthals hunted their competitors to some extent.


title: Neanderthal

text: Neanderthals and cave hyaenas may have exemplified niche differentiation, and actively avoided competing with each other. Although they both mainly targeted the same groups of creatures—deer, horses, and cattle—Neanderthals mainly hunted the former and cave hyaenas the latter two. Further, animal remains from Neanderthal caves indicate they preferred to hunt prime individuals, whereas cave hyaenas hunted weaker or younger prey, and cave hyaena caves have a higher abundance of carnivore remains. Nonetheless, there is evidence that cave hyaenas stole food and leftovers from Neanderthal campsites and scavenged on dead Neanderthal bodies.


title: Neanderthal

text: There are several instances of Neanderthals practising cannibalism across their range. The first example came from Krapina, Croatia, in 1899, and other examples were found at Cueva del Sidrón and Zafarraya in Spain; and the French Grotte de Moula-Guercy, Les Pradelles, and La Quina. For the five cannibalised Neanderthals at the Grottes de Goyet, Belgium, there is evidence that the upper limbs were disarticulated, the lower limbs defleshed and also smashed (likely to extract bone marrow), the chest cavity disemboweled, and the jaw dismembered. There is also evidence that the butchers used some bones to retouch their tools. The processing of Neanderthal meat at Grottes de Goyet is similar to how they processed horse and reindeer. About 35% of the Neanderthals at Marillac-le-Franc, France, show clear signs of butchery, and the presence of digested teeth indicates that the bodies were abandoned and eaten by scavengers, likely hyaenas.


title: Neanderthal

text: These cannibalistic tendencies have been explained as either ritual defleshing, pre-burial defleshing (to prevent scavengers or foul smell), an act of war, or simply for food. Due to a small number of cases, and the higher number of cut marks seen on cannibalised individuals than animals (indicating inexperience), cannibalism was probably not a very common practice, and it may have only been done in times of extreme food shortages as in some cases in recorded human history.


title: Neanderthal

text: Neanderthals used ochre, a clay earth pigment. Ochre is well-documented from 60 to 45 thousand years ago in Neanderthal sites, with the earliest example dating to 250–200 thousand years ago from Maastricht-Belvédère, the Netherlands (a similar timespan to the ochre record of "H. sapiens"). It has been hypothesised to have functioned as body paint, and analyses of pigments from Pech de l’Azé, France, indicates they were applied to soft materials (such as a hide or human skin). However, modern hunter gatherers, in addition to body paint, also use ochre for medicine, for tanning hides, as a food preservative, and as an insect repellent, so its use as decorative paint for Neanderthals is speculative. Containers apparently used for mixing ochre pigments were found in Peștera Cioarei, Romania, which could indicate modification of ochre for solely aesthetic purposes.


title: Neanderthal

text: Neanderthals collected uniquely shaped objects and are suggested to have modified them into pendants, such as a fossil "Aspa marginata" sea snail shell possibly painted red from Grotta di Fumane, Italy, transported over to the site about 47,500 years ago; 3 shells, dated to about 120–115 thousand years ago, perforated through the umbo belonging to a rough cockle, a "Glycymeris insubrica", and a "Spondylus gaederopus" from Cueva de los Aviones, Spain, the former two associated with red and yellow pigments, and the latter a red-to-black mix of hematite and pyrite; and a king scallop shell with traces of an orange mix of goethite and hematite from Cueva Antón, Spain. The discoverers of the latter two claim that pigment was applied to the exterior to make it match the naturally vibrant inside colouration. Excavated from 1949 to 1963 from the French Grotte du Renne, Châtelperronian beads made from animal teeth, shells, and ivory were found associated with Neanderthal bones, but the dating is uncertain and Châtelperronian artefacts may actually have been crafted by modern humans and simply redeposited with Neanderthal remains.