When Yang and Mills proposed the theory, there was no candidate vector boson. J. J. Sakurai in 1960 predicted that there should be a massive vector boson which is coupled to isospin, and predicted that it would show universal couplings. The rho mesons were discovered a short time later, and were quickly identified as Sakurai's vector bosons. The couplings of the rho to the nucleons and to each other were verified to be universal, as best as experiment could measure. The fact that the diagonal isospin current contains part of the electromagnetic current led to the prediction of rho-photon mixing and the concept of vector meson dominance, ideas which led to successful theoretical pictures of GeV-scale photon-nucleus scattering.