After the fall of the Western Roman Empire, astronomy developed further in India and the medieval Islamic world. In 499 CE, the Indian astronomer Aryabhata propounded a planetary model that explicitly incorporated Earth's rotation about its axis, which he explains as the cause of what appears to be an apparent westward motion of the stars. He also theorised that the orbits of planets were elliptical. Aryabhata's followers were particularly strong in South India, where his principles of the diurnal rotation of Earth, among others, were followed and a number of secondary works were based on them.