Big Bang nucleosynthesis is the theory of the formation of the elements in the early universe. It finished when the universe was about three minutes old and its temperature dropped below that at which nuclear fusion could occur. Big Bang nucleosynthesis had a brief period during which it could operate, so only the very lightest elements were produced. Starting from hydrogen ions (protons), it principally produced deuterium, helium-4, and lithium. Other elements were produced in only trace abundances. The basic theory of nucleosynthesis was developed in 1948 by George Gamow, Ralph Asher Alpher, and Robert Herman. It was used for many years as a probe of physics at the time of the Big Bang, as the theory of Big Bang nucleosynthesis connects the abundances of primordial light elements with the features of the early universe. Specifically, it can be used to test the equivalence principle, to probe dark matter, and test neutrino physics. Some cosmologists have proposed that Big Bang nucleosynthesis suggests there is a fourth "sterile" species of neutrino.