In the sentence extracted from biomedical literature, identify all the disease entities. The required answer format is the same sentence with HTML <span> tags to mark up specific entities. 
### Entity Markup Guides:
Use <span class="disease"> to denote a disease.

### Input Text: Chromosomal locations of the Atm ( ataxia - telangiectasia ( AT ) - mutated ) and Acat1 ( mitochondrial acetoacetyl - CoA thiolase ) genes in mouse , rat , and Syrian hamster were determined by direct R - banding FISH .
### Output Text:
Chromosomal locations of the Atm ( <span class="disease">ataxia - telangiectasia</span> ( <span class="disease">AT</span> ) - mutated ) and Acat1 ( mitochondrial acetoacetyl - CoA thiolase ) genes in mouse , rat , and Syrian hamster were determined by direct R - banding FISH . 