In the sentence extracted from biomedical literature, identify all the disease entities. The required answer format is the same sentence with HTML <span> tags to mark up specific entities. 
### Entity Markup Guides:
Use <span class="disease"> to denote a disease.

### Input Text: Comparison of the linkage map of mouse chromosome 9 ( MMU9 ) and that of human chromosome 11 ( HSA11 ) indicates that there is a chromosomal rearrangement due to an inversion between Ets1 and Atm - Npat - Acat1 and that the inversion of MMU9 originated from the chromosomal breakage at the boundary between Gria4 and Atm - Npat - Acat1 on HSA11 .
### Output Text:
Comparison of the linkage map of mouse chromosome 9 ( MMU9 ) and that of human chromosome 11 ( HSA11 ) indicates that there is a chromosomal rearrangement due to an inversion between Ets1 and Atm - Npat - Acat1 and that the inversion of MMU9 originated from the chromosomal breakage at the boundary between Gria4 and Atm - Npat - Acat1 on HSA11 . 