W. D. Hamilton developed the "inclusive" form of this quantity, which explains self-sacrificing behavior.
This quantity names the landscape proposed by Sewall Wright that contains peaks and valleys. The maximum value of this quantity minus the average value all divided by the maximum value equals the genetic load. Fisher's fundamental theorem of natural selection concerns the rate of increase in this quantity. W. D. Hamilton developed the "inclusive" form of this quantity, which explains self-sacrificing behavior. The relative form of this quantity is zero for sterile animals, even if they are robust. For 10 points, name this quantity from evolutionary biology that is the contribution an individual makes to the next generation's gene pool. 
This quantity names the landscape proposed by Sewall Wright that contains peaks and valleys.
fitness [accept specific types] 