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Numerical investigations of this process following Haigh's suggestion assumes the optimal class is an isolated population, and Mukai's observations of viability modifiers on the second chromosome of Drosophila melanogaster can provides estimates for its efficiency. Its effectiveness is dependent on the number of individuals N and thus this model's rate is equal to N times e to the quantity negative U divided by s, where s is the selection coefficient and U is the Poisson mean number. In small populations this process is dominated by the establishment phase and is similar to the "error-catastrophe" theory of somatic aging suggested by Orgel. It has been shown that this model is mathematically equivalent to the Hill-Robertson effect and the current name for this eponymous mechanism is attributed to a 1974 paper by Felsenstein, "The Evolutionary Advantage of Recombination." Its namesake won the 1946 Nobel for his discovery that X-Rays increase the rate of mutation. Proposing to explain the comparative advantage of ?sexual reproduction and co-developed by R.A. Fisher, for 10 points, name this model that describes how genomes of asexual populations irreversibly accumulate deleterious mutations and for which its efficiency is described in number of "turns."
Numerical investigations of this process following Haigh's suggestion assumes the optimal class is an isolated population, and Mukai's observations of viability modifiers on the second chromosome of Drosophila melanogaster can provides estimates for its efficiency.
{Muller's Ratchet} [accept {Fisher}-{Muller Mechanism}]