1. Trees or shrubs; leaves simple; fruits pediceled drupes.


1. Herbs or herbaceous vines; larger leaves compound or lobed; fruits sessile achenes subtended by bracts.

2. Leaves palmately compound with lanceolate to oblanceolate leaflets, opposite on lower portion of the erect stem and alternate above; stems and petioles finely antrorse-hispid; achenes subtended by narrow bracts, not concealed.


2. Leaves at most deeply lobed, all opposite on a climbing stem (vine); stems and petioles retrorse-scabrous; achenes concealed by broad, strongly overlapping bracts.


