Microtubules.
Vesicles Carry Cargo. Most molecules, including proteins, are too large to pass directly through membranes. Instead, large molecules are loaded into small membrane-wrapped containers called vesicles. Vesicles are constantly forming-especially at the plasma membrane, the ER, and the Golgi. Once formed, vesicles deliver their contents to destinations within or outside of the cell. A vesicle forms when the membrane bulges out and pinches off. It travels to its destination then merges with another membrane to release its cargo. In this way proteins and other large molecules are transported without ever having to cross a membrane. Some vesicles form with the help of coat proteins. Geometrically arranged coat proteins on the surface of the membrane help the vesicle to bud off Specialized motor proteins are attached to both the organelle and the microtubules (via chemical interactions). These proteins, called kinesins (which generally move towards the periphery) and dyneins (which move towards the center) walk the organelle up and down the microtubules. (picture to the left, from Molecular Probes) This cytoskeleton is the surface that organelles can move on. Among the proteins that make up the cytoskeleton, microtubules are the predominant walkway organelles use to move around the cell. Cell Parts and Organelle Functions -. The cell membrane (or plasma Membrane) is a structure that forms the outer boundary of the cell and allows only certain materials to move into and out of the cell. Food, oxygen and water move into the cell through the membrane. Waste products also leave through the membrane. In cells, structures called Golgi Bodies (also known as Golgi apparatus or Golgi complex) are stacks of membrane-covered sacs that package and move proteins to the outside of the cell. Golgi bodies are the packaging and secreting organelles 1 Organelle movement. 2  Microtubules and microfilaments can help move organelles from place to place in the cell. 3  In the case of endocytosis when a vesicle is formed to engulf some particle from outside the cell, microfilaments attach to the vesicle and pull it into the cell. Most eukaryotic cells contain a complex network of protein fibers called the cytoskeleton. It forms a framework for the movement of organelles around the cytoplasm-most of the organelles are attached to the cytoskeleton. The network consists of protein microfilaments, intermediate filaments, and microtubules. The cytoskeleton provides an important structural framework for: 1  Cell shape. 2  For cells without cell walls, the cytoskeleton determines the shape of the cell. 3  This is one of the functions of the intermediate filaments The membrane enclosing the vesicle is also a lamellar phase, similar to that of the plasma membrane, and vesicles can fuse with the plasma membrane to release their contents outside of the cell. Vesicles can also fuse with other organelles within the cell. Vesicles perform a variety of functions. Vesicles can form naturally, for example, during the processes of secretion (exocytosis), uptake (phagocytosis and endocytosis) and transport of materials within the cytoplasm. Alternatively, they may be prepared artificially, in which case they are called liposome We continue our examination of the cytoskeleton with a look at the role of microtubules in intracellular transport and positioning of vesicles and organelles. We will then move on to look at actin filaments and their functions. In this way, they can guide the movement of materials through the cell (note that actin filaments, which we will discuss later, may also function in this manner). In addition to moving vesicles and organelles microtubules function in chromosome movement during cell division, as we have already seen. Axoplasmic transport, also called axonal transport, is a cellular process responsible for movement of mitochondria, lipids, synaptic vesicles, proteins, and other cell parts (i.e. organelles) to and from a neuron's cell body, through the cytoplasm of its axon (the axoplasm). Retrograde transport is movement of molecules/organelles inward, away from the synapse or plasma membrane toward the cell body or soma. Retrograde transport is mediated by dynein, and is used for example to send chemical messages and endocytosis products headed to endolysosomes from the axon back to the cell. While peroxisomes look similar to lysosomes, they are generally bigger than lysosomes. Peroxisomes are found near mitochondria and chloroplasts, while lysosomes can be found anywhere in the cell. The cytoskeleton is the structure that gives a cell shape, holds organelles in place, and lets parts of the cell move. 
http://learn.genetics.utah.edu/content/cells/vesicles/ http://www.madsci.org/posts/archives/2001-01/980869118.Cb.r.html http://salem.k12.va.us/staff/jwright/CellPartsandOrganelleFunctions.htm http://hyperphysics.phy-astr.gsu.edu/hbase/Biology/cytoskel.html https://en.wikipedia.org/wiki/Vesicle_(biology_and_chemistry) http://oregonstate.edu/instruction/bi314/summer08/cytotwo.html https://en.wikipedia.org/wiki/Axoplasmic_transport http://library.vcc.ca/learningcentre/pdf/vcclc/ReviewOfTheCellAndItsOrganelles.pdf
what moves vesicles and organelles throughout the organelle