If the tone is immediately before the puff, then the learning can be done unconsciously (and in rabbits, done with much of the brain removed).
उनीहरू अलि ढिला भएका कारण उद्धार कार्यमा पठाउन सकिएन तर पञ्चायत कालमा जस्तै उपेक्षा गर्ने मन पनि भएन ।