Under his direction, which continued over a period of 45 years to the end of his life, this Institute became one of the most important centers of physiological research.
تحت نظارت او، که طی یک دوره ۴۵ ساله تا پایان عمر خود ادامه یافت، این موسسه یکی از مهمترین مراکز تحقیق فیزیولوژیک شد.