extinct genus ants subfamily described species described species denmark fossil beds british_columbia canada fourth fossil site russia queens genus large elongated eyes developed present behaviour ants similar extant ants solitary foraging prey leaving trails poor size birds animals likely preyed ants assigned tribe instead generally regarded authors believe assigned history classification fossils studied described cambridge massachusetts published description genus species annals journal article genus combination referring age specimens greek meaning minimum fossil age approximately million years genus description paper contained descriptions type species fossil beds british_columbia highlands lake system cover included danish species new combination fourth species described colleagues originally classified latin uncertain placement subfamily ants identified tribe level paper switzerland expressed doubt members ants notes development area area compound eyes different reduced reduced key traits notes antennae shape type species properly determined making true placement antennae elongated means fossil definitely feature key trait believed classified family subsequent report russian describing new accepted classification colleagues mentioning comments following generated colleagues possible phylogenetic position ants subfamily genus closely related extinct genera including extant description characteristics separate genera waist connecting abdomen composed single segment head species varies shape distinct triangular shape shorter head capsule teeth elongated queens genus large measuring millimetres characters include large developed eyes rounded noticeable described single counterpart compression fossil found near british_columbia type specimens numbered counterpart currently preserved collections housed british_columbia cover coined specific epithet latin meaning rounded circular reference shape head species species notably rounded head capsule shape smoothly sloping convex shape center similar appearance round queen estimated approximately millimetres compound eyes large oval shape antennae described poor preservation large containing seven teeth developed present second species described like known single queen specimen counterpart included ottawa collections shape head separates estimated millimetres smaller shape distinct species specific epithet chosen honor collected specimen honor work antennae preserved eyes distinguished fewer teeth queen preserved portions legs preserved portions deformed specimen appears crushed overall morphology species placement worker form genus possibly confirmed poor preservation species known specimens collected earliest found rocks specimens studied described named species collected type specimen september queen described preserved legs missing counterpart deposited time placed species modern genus based shape abdominal segment lack cover moved species based similarity species shape abdominal segment iii differs found subfamily genera morphology distinguished species shape size head notably larger ants somewhat angular head different species having rounded head species known exclusively queens known male assigned average length queen millimetres robust body head round width length measuring millimetres eyes oval shaped developed located near middle head triangular long teeth present antennae long segments millimetres long millimetres long millimetres wide convex short millimetres long millimetres wide swollen early process transition decaying organism time developed present specimens single complete male known collected specimens body length shorter comparison queen measuring millimetres slender head small eyes fully developed large antennae preserved small fragments legs present smaller elongated preserved described counterpart solitary compression fossil found late fossil site area far eastern russia type specimens numbered pin counterpart currently preserved fossil collections coined specific epithet latin meaning eastern reference type locality species smaller size distinct differences highlands species fossils placed new species notable time difference highlands possible incomplete estimated body length robust fore wing closed cells elongated twice long wide ecology life habits similar extant ants colonies soil trees making nesting species workers likely solitary foraging ground low vegetation trees consuming workers likely recruit lead food sources workers lay trails ants likely large eyes find prey navigational purposes abundance queens collected suggests ants poor large size morphology indicate specialised tools chambers soil wood colony foundation large size common occurrence geographical range mean important food source variety birds animals fed insects
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